blank Quick help
blank Maintenance news

Scheduled maintenance

Regular maintenance outages:
between 05.00 and 05.15 hrs CET (Monday to Sunday).

Other outages
Availability
Register Forum

2022.02.11

More...
blank News flashes

News flashes

New version of the European Patent Register - SPC information for Unitary Patents.

2024-03-06

More...
blank Related links

Release notes
European Patent Register Assistant
European Patent Register Resource Book
European Patent Register – pocket guide
FAQ - European Patent Register
FAQ - Register Alert
Federated Register (Flyer)
Global Dossier (Flyer)
Register events
Deep link coverage
Federated Register content
The Patent Information Tour
Fair use charter

Extract from the Register of European Patents

EP About this file: EP2816105

EP2816105 - GREEN SULFUR BACTERIA VARIANT AND BACTERIOCHLOROPHYLL [Right-click to bookmark this link]
StatusNo opposition filed within time limit
Status updated on  08.02.2019
Database last updated on 06.07.2024
FormerThe patent has been granted
Status updated on  02.03.2018
FormerGrant of patent is intended
Status updated on  04.10.2017
FormerExamination is in progress
Status updated on  25.11.2016
Most recent event   Tooltip01.07.2022Lapse of the patent in a contracting state
New state(s): MK		published on 03.08.2022  [2022/31]
Applicant(s)For all designated states
Kurume University
67, Asahi-machi Kurume-shi
Fukuoka 830-0011 / JP
[2014/52]
Inventor(s)01 / HARADA, Jiro
c/o Kurume University
67 Asahi-machi
Kurume-shi Fukuoka 830-0011 / JP
02 / NOGUCHI, Masato
c/o Kurume University
67 Asahi-machi
Kurume-shi Fukuoka 830-0011 / JP
03 / TAMIAKI, Hitoshi
c/o Department of Pharmacy
College of Pharmaceutical Sciences
Ritsumeikan University
1-1-1 Nojihigashi
Kusatsu-shi Shiga 525-8577 / JP
 [2014/52]
Representative(s)J A Kemp LLP
80 Turnmill Street
London EC1M 5QU / GB
[N/P]
Former [2018/14]J A Kemp
14 South Square
Gray's Inn
London WC1R 5JJ / GB
Former [2014/52]Campbell, Patrick John Henry
J A Kemp
14 South Square
Gray's Inn
London WC1R 5JJ / GB
Application number, filing date13748805.212.02.2013
[2018/14]
WO2013JP53295
Priority number, dateJP2012002891913.02.2012         Original published format: JP 2012028919
[2014/52]
Filing languageJA
Procedural languageEN
PublicationType: A1 Application with search report
No.:WO2013122064
Date:22.08.2013
Language:JA
[2013/34]
Type: A1 Application with search report 
No.:EP2816105
Date:24.12.2014
Language:EN
[2014/52]
Type: B1 Patent specification 
No.:EP2816105
Date:04.04.2018
Language:EN
[2018/14]
Search report(s)International search report - published on:JP22.08.2013
(Supplementary) European search report - dispatched on:EP16.09.2015
ClassificationIPC:C12N1/20, C12N1/21, C12N15/09, C12N9/10, C12P9/00, C12P17/18, C07D487/22, // C12R1/01
[2015/42]
CPC:
C07D487/22 (EP,US); C12N1/205 (EP,US); C12N9/1007 (EP,US);
C12P17/182 (EP,US); C12R2001/01 (EP,US); Y02P20/52 (US)
Former IPC [2014/52]C12N1/20, C12N1/21, C12P9/00, C12N15/09
Designated contracting statesAL,   AT,   BE,   BG,   CH,   CY,   CZ,   DE,   DK,   EE,   ES,   FI,   FR,   GB,   GR,   HR,   HU,   IE,   IS,   IT,   LI,   LT,   LU,   LV,   MC,   MK,   MT,   NL,   NO,   PL,   PT,   RO,   RS,   SE,   SI,   SK,   SM,   TR [2014/52]
Extension statesBANot yet paid
MENot yet paid
TitleGerman:GRÜNE SCHWEFELBAKTERIENVARIANTEN UND BAKTERIOCHLOROPHYLL[2014/52]
English:GREEN SULFUR BACTERIA VARIANT AND BACTERIOCHLOROPHYLL[2014/52]
French:VARIANT DE BACTÉRIES SULFUREUSES VERTES ET BACTÉRIOCHOROPHYLLE[2014/52]
Biological materialThis application mentions deposited biological material, check the file for details
Entry into regional phase12.09.2014Translation filed 
12.09.2014National basic fee paid 
12.09.2014Search fee paid 
12.09.2014Designation fee(s) paid 
12.09.2014Examination fee paid 
Examination procedure12.09.2014Examination requested  [2014/52]
04.04.2016Amendment by applicant (claims and/or description)
25.11.2016Despatch of a communication from the examining division (Time limit: M04)
29.03.2017Reply to a communication from the examining division
05.10.2017Communication of intention to grant the patent
24.01.2018Fee for grant paid
24.01.2018Fee for publishing/printing paid
24.01.2018Receipt of the translation of the claim(s)
Divisional application(s)The date of the Examining Division's first communication in respect of the earliest application for which a communication has been issued is  25.11.2016
Opposition(s)07.01.2019No opposition filed within time limit [2019/11]
Fees paidRenewal fee
11.02.2015Renewal fee patent year 03
10.02.2016Renewal fee patent year 04
10.02.2017Renewal fee patent year 05
26.02.2018Renewal fee patent year 06
Opt-out from the exclusive  Tooltip
competence of the Unified
Patent Court		 See the Register of the Unified Patent Court for opt-out data
Responsibility for the accuracy, completeness or quality of the data displayed under the link provided lies entirely with the Unified Patent Court.
Lapses during opposition  TooltipHU12.02.2013
AL04.04.2018
AT04.04.2018
CY04.04.2018
CZ04.04.2018
DK04.04.2018
EE04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
IT04.04.2018
LT04.04.2018
LV04.04.2018
MC04.04.2018
MK04.04.2018
NL04.04.2018
PL04.04.2018
RO04.04.2018
RS04.04.2018
SE04.04.2018
SI04.04.2018
SK04.04.2018
SM04.04.2018
TR04.04.2018
BG04.07.2018
NO04.07.2018
GR05.07.2018
IS04.08.2018
PT06.08.2018
[2022/31]
Former [2021/32]HU12.02.2013
AL04.04.2018
AT04.04.2018
CY04.04.2018
CZ04.04.2018
DK04.04.2018
EE04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
IT04.04.2018
LT04.04.2018
LV04.04.2018
MC04.04.2018
NL04.04.2018
PL04.04.2018
RO04.04.2018
RS04.04.2018
SE04.04.2018
SI04.04.2018
SK04.04.2018
SM04.04.2018
TR04.04.2018
BG04.07.2018
NO04.07.2018
GR05.07.2018
IS04.08.2018
PT06.08.2018
Former [2021/31]AL04.04.2018
AT04.04.2018
CY04.04.2018
CZ04.04.2018
DK04.04.2018
EE04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
IT04.04.2018
LT04.04.2018
LV04.04.2018
MC04.04.2018
NL04.04.2018
PL04.04.2018
RO04.04.2018
RS04.04.2018
SE04.04.2018
SI04.04.2018
SK04.04.2018
SM04.04.2018
TR04.04.2018
BG04.07.2018
NO04.07.2018
GR05.07.2018
IS04.08.2018
PT06.08.2018
Former [2021/26]AL04.04.2018
AT04.04.2018
CY04.04.2018
CZ04.04.2018
DK04.04.2018
EE04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
IT04.04.2018
LT04.04.2018
LV04.04.2018
MC04.04.2018
NL04.04.2018
PL04.04.2018
RO04.04.2018
RS04.04.2018
SE04.04.2018
SI04.04.2018
SK04.04.2018
SM04.04.2018
TR04.04.2018
BG04.07.2018
NO04.07.2018
GR05.07.2018
PT06.08.2018
Former [2020/16]AL04.04.2018
AT04.04.2018
CZ04.04.2018
DK04.04.2018
EE04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
IT04.04.2018
LT04.04.2018
LV04.04.2018
MC04.04.2018
NL04.04.2018
PL04.04.2018
RO04.04.2018
RS04.04.2018
SE04.04.2018
SI04.04.2018
SK04.04.2018
SM04.04.2018
TR04.04.2018
BG04.07.2018
NO04.07.2018
GR05.07.2018
PT06.08.2018
Former [2019/45]AL04.04.2018
AT04.04.2018
CZ04.04.2018
DK04.04.2018
EE04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
IT04.04.2018
LT04.04.2018
LV04.04.2018
MC04.04.2018
NL04.04.2018
PL04.04.2018
RO04.04.2018
RS04.04.2018
SE04.04.2018
SI04.04.2018
SK04.04.2018
SM04.04.2018
BG04.07.2018
NO04.07.2018
GR05.07.2018
PT06.08.2018
Former [2019/26]AL04.04.2018
AT04.04.2018
CZ04.04.2018
DK04.04.2018
EE04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
IT04.04.2018
LT04.04.2018
LV04.04.2018
NL04.04.2018
PL04.04.2018
RO04.04.2018
RS04.04.2018
SE04.04.2018
SI04.04.2018
SK04.04.2018
SM04.04.2018
BG04.07.2018
NO04.07.2018
GR05.07.2018
PT06.08.2018
Former [2019/11]AL04.04.2018
AT04.04.2018
CZ04.04.2018
DK04.04.2018
EE04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
IT04.04.2018
LT04.04.2018
LV04.04.2018
NL04.04.2018
PL04.04.2018
RO04.04.2018
RS04.04.2018
SE04.04.2018
SK04.04.2018
SM04.04.2018
BG04.07.2018
NO04.07.2018
GR05.07.2018
PT06.08.2018
Former [2019/10]AL04.04.2018
AT04.04.2018
CZ04.04.2018
DK04.04.2018
EE04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
LT04.04.2018
LV04.04.2018
NL04.04.2018
PL04.04.2018
RO04.04.2018
RS04.04.2018
SE04.04.2018
SK04.04.2018
BG04.07.2018
NO04.07.2018
GR05.07.2018
PT06.08.2018
Former [2019/09]AL04.04.2018
DK04.04.2018
EE04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
LT04.04.2018
LV04.04.2018
NL04.04.2018
PL04.04.2018
RS04.04.2018
SE04.04.2018
BG04.07.2018
NO04.07.2018
GR05.07.2018
PT06.08.2018
Former [2019/08]AL04.04.2018
DK04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
LT04.04.2018
LV04.04.2018
NL04.04.2018
PL04.04.2018
RS04.04.2018
SE04.04.2018
BG04.07.2018
NO04.07.2018
GR05.07.2018
PT06.08.2018
Former [2019/03]AL04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
LT04.04.2018
LV04.04.2018
NL04.04.2018
PL04.04.2018
RS04.04.2018
SE04.04.2018
BG04.07.2018
NO04.07.2018
GR05.07.2018
PT06.08.2018
Former [2018/52]AL04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
LT04.04.2018
LV04.04.2018
NL04.04.2018
PL04.04.2018
RS04.04.2018
SE04.04.2018
BG04.07.2018
NO04.07.2018
GR05.07.2018
Former [2018/51]AL04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
LT04.04.2018
LV04.04.2018
NL04.04.2018
PL04.04.2018
RS04.04.2018
SE04.04.2018
BG04.07.2018
NO04.07.2018
Former [2018/50]AL04.04.2018
ES04.04.2018
FI04.04.2018
HR04.04.2018
LT04.04.2018
NL04.04.2018
PL04.04.2018
SE04.04.2018
BG04.07.2018
NO04.07.2018
Former [2018/49]AL04.04.2018
ES04.04.2018
FI04.04.2018
LT04.04.2018
NL04.04.2018
PL04.04.2018
SE04.04.2018
BG04.07.2018
NO04.07.2018
Former [2018/47]AL04.04.2018
ES04.04.2018
FI04.04.2018
LT04.04.2018
NL04.04.2018
NO04.07.2018
Former [2018/46]AL04.04.2018
LT04.04.2018
NL04.04.2018
NO04.07.2018
Former [2018/43]NL04.04.2018
Documents cited:Search[IY]  - MIYATAKE T ET AL, "Self-aggregates of bacteriochlorophylls-c, d and e in a light-harvesting antenna system of green photosynthetic bacteria: Effect of stereochemistry at the chiral 3-(1-hydroxyethyl) group on the supramolecular arrangement of chlorophyllous pigments", JOURNAL OF PHOTOCHEMISTRY AND PHOTOBIOLOGY C: PHOTOCHEMISTRY REVIEWS, (20050801), vol. 6, no. 2-3, ISSN 1389-5567, pages 89 - 107, XP027707342 [I] 6-9 * 4. Self-aggregates of modified unnatural BChls possessing a chiral secondary alcoholic group * [Y] 10
 [Y]  - SEIJI TOKITA ET AL, "A novel and mild isolation procedure of chlorosomes from the green sulfur bacterium Chlorobaculum tepidum", PHOTOSYNTHESIS RESEARCH, (20110826), vol. 108, no. 2-3, doi:10.1007/s11120-011-9679-4, ISSN 0166-8595, pages 183 - 190, XP055211138 [Y] 10 * the whole document *

DOI:   http://dx.doi.org/10.1007/s11120-011-9679-4
 [Y]  - OOSTERGETEL ET AL, "Long-range organization of bacteriochlorophyll in chlorosomes of Chlorobium tepidum investigated by cryo-electron microscopy", FEBS LETTERS, (20071105), vol. 581, no. 28, ISSN 0014-5793, pages 5435 - 5439, XP022374807 [Y] 10 * the whole document *
 [A]  - J. A. MARESCA ET AL, "The bchU Gene of Chlorobium tepidum Encodes the C-20 Methyltransferase in Bacteriochlorophyll c Biosynthesis", JOURNAL OF BACTERIOLOGY, (20040416), vol. 186, no. 9, doi:10.1128/JB.186.9.2558-2566.2004, ISSN 0021-9193, pages 2558 - 2566, XP055210909 [A] 2-7 * the whole document *

DOI:   http://dx.doi.org/10.1128/JB.186.9.2558-2566.2004
 [A]  - HARADA J ET AL, "In vitro activity of C-20 methyltransferase, BchU, involved in bacteriochlorophyll c biosynthetic pathway in green sulfur bacteria", FEBS LETTERS, vol. 579, no. 9, ISSN 0014-5793, (20050302), pages 1983 - 1987, (20050328), XP027696485 [A] 2-7 * the whole document *
 [T]  - YUSUKE TSUKATANI ET AL, "Bacteriochlorophyll homolog compositions in the bchU mutants of green sulfur bacteria", PHOTOCHEMICAL & PHOTOBIOLOGICAL SCIENCES, (20131001), vol. 12, no. 12, doi:10.1039/c3pp50253h, ISSN 1474-905X, page 2195, XP055210879 [T] 1-10 * the whole document *

DOI:   http://dx.doi.org/10.1039/c3pp50253h
 [T]  - ORF GREGORY S ET AL, "Chlorosome antenna complexes from green photosynthetic bacteria", PHOTOSYNTHESIS RESEARCH, (20130613), vol. 116, no. 2, doi:10.1007/S11120-013-9869-3, ISSN 0166-8595, pages 315 - 331, XP035355276 [T] 1-10 * the whole document *

DOI:   http://dx.doi.org/10.1007/s11120-013-9869-3
 [T]  - DARIUSZ M. NIEDZWIEDZKI ET AL, "Photophysical Properties of the Excited States of Bacteriochlorophyll f in Solvents and in Chlorosomes", THE JOURNAL OF PHYSICAL CHEMISTRY B, (20140110), vol. 118, no. 9, doi:10.1021/jp409495m, ISSN 1520-6106, pages 2295 - 2305, XP055210899 [T] 1-10 * the whole document *

DOI:   http://dx.doi.org/10.1021/jp409495m
International search[A]  - HITOSHI TAMIAKI ET AL., "Synthesis of homologously pure bacteriochlorophyll-e and f analogues from BChls-c/d via transformation of the 7-methyl to formyl group and self- aggregation of synthetic zinc methyl bacteriopheophorbides-c/d/e/f in non-polar organic solvent.", TETRAHEDRON, (2003), vol. 59, pages 4337 - 4350, XP004429347

DOI:   http://dx.doi.org/10.1016/S0040-4020(03)00638-0
 [A]  - NIKOLAUS RISCH, "Bacteriochlorophyll f. - Partial synthesis and the behavior of some derivatives.", LIEBIGS ANNALEN DER CHEMIE, (1988), vol. 4, pages 343 - 347, XP055164831

DOI:   http://dx.doi.org/10.1002/jlac.198819880407
 [A]  - HITOSHI TAMIAKI ET AL., "A novel approach toward bacteriochlorophylls-e and f.", BIOORGANIC & MEDICINAL CHEMISTRY LETTERS, (1999), vol. 9, pages 1631 - 1632, XP004167729

DOI:   http://dx.doi.org/10.1016/S0960-894X(99)00258-9
 [A]  - SHIN-ICHI SASAKI ET AL., "Self- assembly of synthetic bacteriochlorophyll-f analogues having C8-formyl group", BULLETIN OF THE CHEMICAL SOCIETY OF JAPAN, (2004), vol. 77, pages 797 - 800, XP008174498

DOI:   http://dx.doi.org/10.1246/bcsj.77.797
 [A]  - MIN CHEN ET AL., "A red-shifted chlorophyll.", SCIENCE, (2010), vol. 329, pages 1318 - 1319, XP055161830

DOI:   http://dx.doi.org/10.1126/science.1191127
 [A]  - HITOSHI TAMIAKI ET AL., "In vitro synthesis and characterization of bacteriochlorophyll-f and its absence in bacteriochlorophyll-e producing organisms.", PHOTOSYNTHESIS RESEARCH, (2011), vol. 107, pages 133 - 138, XP019880444

DOI:   http://dx.doi.org/10.1007/s11120-010-9603-3
 [A]  - HITOSHI TAMIAKI ET AL., "Analysis of Bacteriochlorophyl", LOW TEMPERATURE SCIENCE, (2009), vol. 67, pages 339 - 346, XP008175005
 [XP]  - JIRO HARADA ET AL., "A seventh bacterial chlorophyll driving a large light-harvesting antenna.", SCIENTIFIC REPORTS, (201209), vol. 2, page 671, XP055161838

DOI:   http://dx.doi.org/10.1038/srep00671
 [XP]  - KAJETAN VOGL ET AL., "Bacteriochlorophyll f: properties of chlorosomes containing the ''forbidden chlorophyll''.", FRONTIERS IN MICROBIOLOGY, (201208), vol. 3, page 298, XP055161840

DOI:   http://dx.doi.org/10.3389/fmicb.2012.00298
 [XP]  - GREGORY S. ORF ET AL., "Spectroscopic insights into the decreased efficiency of chlorosomes containing bacteriochlorophyll f.", BIOCHIMICA ET BIOPHYSICA ACTA, (20130124), vol. 1827, pages 493 - 501, XP055161841

DOI:   http://dx.doi.org/10.1016/j.bbabio.2013.01.006
by applicant   - H. TAMIAKI, COORD. CHEM. REV., (1996), vol. 148, pages 183 - 197
    - V. I. PROKHORENKO ET AL., J. PHYS. CHEM. B, (2002), vol. 106, pages 5761 - 5768
    - N. U. FRIGAARD ET AL., PHOTOSYNTH. RES., (2003), vol. 78, pages 93 - 117
    - N. U. FRIGAARD, METHODS MOL. BIOL., (2004), vol. 274, pages 325 - 340
    - S. GANAPATHY ET AL., PROC. NATL. ACAD. SCI. USA., (2009), vol. 106, pages 8525 - 8530
    - FRIGAAD NU; BRYANT DA, APPL. ENVIRON. MICROBIOL., (2001), vol. 67, pages 2538 - 2544
    - FRIGAAD NU; SAKURAGI Y; BRYANT DA, METHODS OL. BIOL., (2004), vol. 274, pages 325 - 340
    - ENVIRON. MICROBIOL., (2001), vol. 67, pages 2538 - 2544
    - METHODS MOL. BIOL., (2004), vol. 274, pages 325 - 340
    - H. TAMIAKI ET AL., PHOTOSYNTH. RES., (2011), vol. 107, pages 133 - 138
The EPO accepts no responsibility for the accuracy of data originating from other authorities; in particular, it does not guarantee that it is complete, up to date or fit for specific purposes.