EP3174978 - PROTEIN WITH DEXTRAN-SACCHARASE ACTIVITY, AND USES [Right-click to bookmark this link] | Status | No opposition filed within time limit Status updated on 17.07.2020 Database last updated on 03.09.2024 | |
Former | The patent has been granted Status updated on 09.08.2019 | ||
Former | Grant of patent is intended Status updated on 13.05.2019 | ||
Former | Examination is in progress Status updated on 08.12.2017 | ||
Former | Request for examination was made Status updated on 15.06.2017 | ||
Former | The international publication has been made Status updated on 13.12.2016 | Most recent event Tooltip | 08.07.2022 | Lapse of the patent in a contracting state New state(s): MK | published on 10.08.2022 [2022/32] | Applicant(s) | For all designated states Institut National des Sciences Appliquées de Toulouse 135 avenue de Rangueil 31077 Toulouse Cedex 4 / FR | For all designated states Institut national de recherche pour l'agriculture, l'alimentation et l'environnement 147 Rue de l'Université 75007 Paris / FR | For all designated states Centre National de la Recherche Scientifique 3, rue Michel-Ange 75016 Paris / FR | For all designated states Université Paul Sabatier Toulouse III 118 route de Narbonne 31400 Toulouse / FR | [N/P] |
Former [2020/24] | For all designated states Institut National des Sciences Appliquées de Toulouse 135 avenue de Rangueil 31077 Toulouse Cedex 4 / FR | ||
For all designated states Institut national de recherche pour l'agriculture, l'alimentation et l'environnement 147 rue de l'Université 75007 Paris / FR | |||
For all designated states Centre National de la Recherche Scientifique 3, rue Michel-Ange 75016 Paris / FR | |||
For all designated states Université Paul Sabatier Toulouse III 118 route de Narbonne 31400 Toulouse / FR | |||
Former [2017/23] | For all designated states Institut National des Sciences Appliquées de Toulouse 135 avenue de Rangueil 31077 Toulouse Cedex 4 / FR | ||
For all designated states Institut National De La Recherche Agronomique 147, rue de l'Université 75007 Paris / FR | |||
For all designated states Centre National de la Recherche Scientifique 3, rue Michel-Ange 75016 Paris / FR | |||
For all designated states Université Paul Sabatier Toulouse III 118 route de Narbonne 31400 Toulouse / FR | Inventor(s) | 01 /
VUILLEMIN, Marlène Heslegårdsvej 8, 3-4 2900 Hellerup / DK | 02 /
CLAVERIE, Marion 4 Rue Sully F-64800 Benejacq / FR | 03 /
MOULIS, Claire Lazières F-31290 Viellevigne / FR | 04 /
REMAUD-SIMEON, Magali 1 Rue Benjamin Charrié F-31520 Ramonville-Saint-Agne / FR | 05 /
MONSAN, Pierre 22 Chemin de la Gravette F-31700 Mondonville / FR | 06 /
SEVERAC, Etienne 13 Avenue de Gasgogne F-31820 Pibrac / FR | 07 /
FONTAGNE-FAUCHER, Catherine L'Haouré Chemin de Beaupuy F-32600 L'Isle Jourdain / FR | [2017/23] | Representative(s) | Plasseraud IP 104 Rue de Richelieu CS92104 75080 Paris Cedex 02 / FR | [N/P] |
Former [2019/37] | Cabinet Plasseraud 66, rue de la Chaussée d'Antin 75440 Paris Cedex 09 / FR | ||
Former [2017/23] | Cabinet Plasseraud 66 rue de la Chaussée d'Antin 75440 Paris Cedex 09 / FR | Application number, filing date | 15759857.4 | 21.07.2015 | [2017/23] | WO2015FR52002 | Priority number, date | FR20140057264 | 28.07.2014 Original published format: FR 1457264 | [2017/23] | Filing language | FR | Procedural language | FR | Publication | Type: | A1 Application with search report | No.: | WO2016016544 | Date: | 04.02.2016 | Language: | FR | [2016/05] | Type: | A1 Application with search report | No.: | EP3174978 | Date: | 07.06.2017 | Language: | FR | The application published by WIPO in one of the EPO official languages on 04.02.2016 takes the place of the publication of the European patent application. | [2017/23] | Type: | B1 Patent specification | No.: | EP3174978 | Date: | 11.09.2019 | Language: | FR | [2019/37] | Search report(s) | International search report - published on: | EP | 04.02.2016 | Classification | IPC: | C12N9/10, C12P19/08, C12P19/18, A61K8/73, A61K47/36, A61K31/721, C08B37/02, A23L29/269 | [2019/20] | CPC: |
C12N9/1051 (EP,US);
A23L29/273 (EP,US);
A61K31/721 (EP,US);
A61K47/36 (US);
A61K8/73 (US);
C08B37/0021 (US);
C12P19/08 (EP,US);
C12P19/18 (EP,US);
C12Y204/01005 (EP,US);
A23V2002/00 (US)
(-)
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Former IPC [2017/23] | C12N9/10, C12P19/08, C12P19/18 | Designated contracting states | AL, AT, BE, BG, CH, CY, CZ, DE, DK, EE, ES, FI, FR, GB, GR, HR, HU, IE, IS, IT, LI, LT, LU, LV, MC, MK, MT, NL, NO, PL, PT, RO, RS, SE, SI, SK, SM, TR [2017/23] | Extension states | BA | Not yet paid | ME | Not yet paid | Validation states | MA | Not yet paid | Title | German: | PROTEIN MIT DEXTRANSACCHARASEAKTIVITÄT UND VERWENDUNGEN | [2017/23] | English: | PROTEIN WITH DEXTRAN-SACCHARASE ACTIVITY, AND USES | [2017/23] | French: | PROTEINE A ACTIVITE DEXTRANE-SACCHARASE ET APPLICATIONS | [2017/23] | Entry into regional phase | 13.12.2016 | National basic fee paid | 13.12.2016 | Designation fee(s) paid | 13.12.2016 | Examination fee paid | Examination procedure | 13.12.2016 | Amendment by applicant (claims and/or description) | 13.12.2016 | Examination requested [2017/23] | 13.12.2016 | Date on which the examining division has become responsible | 12.12.2017 | Despatch of a communication from the examining division (Time limit: M06) | 11.06.2018 | Reply to a communication from the examining division | 07.08.2018 | Despatch of a communication from the examining division (Time limit: M06) | 03.01.2019 | Reply to a communication from the examining division | 14.05.2019 | Communication of intention to grant the patent | 07.06.2019 | Fee for grant paid | 07.06.2019 | Fee for publishing/printing paid | 07.06.2019 | Receipt of the translation of the claim(s) | Opposition(s) | 15.06.2020 | No opposition filed within time limit [2020/34] | Fees paid | Renewal fee | 24.05.2017 | Renewal fee patent year 03 | 27.07.2018 | Renewal fee patent year 04 | 30.07.2019 | Renewal fee patent year 05 |
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Responsibility for the accuracy, completeness or quality of the data displayed under the link provided lies entirely with the Unified Patent Court. | Lapses during opposition Tooltip | AL | 11.09.2019 | AT | 11.09.2019 | CY | 11.09.2019 | CZ | 11.09.2019 | DK | 11.09.2019 | EE | 11.09.2019 | FI | 11.09.2019 | HR | 11.09.2019 | LT | 11.09.2019 | LV | 11.09.2019 | MC | 11.09.2019 | MK | 11.09.2019 | MT | 11.09.2019 | PL | 11.09.2019 | RO | 11.09.2019 | RS | 11.09.2019 | SI | 11.09.2019 | SK | 11.09.2019 | SM | 11.09.2019 | TR | 11.09.2019 | BG | 11.12.2019 | GR | 12.12.2019 | IS | 12.01.2020 | [2022/32] |
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Former [2020/08] | LT | 11.09.2019 | Cited in | International search | [A]EP0164656 (PFEIFER & LANGEN [DE]) [A] 11,12 * claims 1-19 *; | [Y]US4767614 (SCARPA IOANNIS S [US], et al) [Y] 10,13 * example 1 *; | [A]US5229277 (DAY DONAL F [US], et al) [A] 10,13 * the whole document *; | [Y]WO2010129839 (TATE & LYLE INGREDIENTS FRANCE [FR], et al) [Y] 10,13 * example 4 *; | [A]WO2010128859 (UNIV GRONINGEN [NL], et al) [A] 11,12 * page 9 - page 12 *; | [Y] - DOLS M ET AL, "Characterization of the different dextransucrase activities excreted in glucose, fructose, or sucrose medium by Leuconostoc mesenteroides NRRL B-1299", APPLIED AND ENVIRONMENTAL MICROBIOLOGY, AMERICAN SOCIETY FOR MICROBIOLOGY, US, (199804), vol. 64, no. 4, ISSN 0099-2240, pages 1298 - 1302, XP002360116 [Y] 10,13 * page 1299 - page 1301 * DOI: http://dx.doi.org/10.1128/AEM.64.4.1298-1302.1998 | [Y] - SIMPSON C ET AL, "Four glucosyltransferases, GtfJ, GtfK, GtfL, and GtfM from Streptococcus salivarius ATCC 25975", MICROBIOLOGY AND IMMUNOLOGY, CENTER FOR ACADEMIC PUBLICATIONS JAPAN|, JP, (199506), vol. 141, no. 6, ISSN 0385-5600, pages 1451 - 1460, XP002082272 [Y] 10,13 * table 2 * | [Y] - Anonymous, "Dextrans", (20140120), URL: https://web.archive.org/web/20140120040848/http://www.sigmaaldrich.com/life-science/biochemicals/biochemical-products.html?TablePage=22696471, (20151005), XP055218107 [Y] 10,13 * page 1 - page 2 * | [Y] - Vicki Caligur, "Dextran and Related Polysaccharides", BioFiles, (2008), URL: http://www.sigmaaldrich.com/content/dam/sigma-aldrich/articles/biofiles/volume3.10a17/Dextran-and-Related-Polysaccharides.pdf, (20151005), XP055218106 [Y] 10,13 * page 1 - page 2 * | [Y] - ELVIRA KHALIKOVA ET AL, "Microbial Dextran-Hydrolyzing Enzymes: Fundamentals and Applications", MICROBIOLOGY AND MOLECULAR BIOLOGY REVIEWS, (200506), vol. 69, no. 2, doi:10.1128/JMBR.69.2.306-325.2005, pages 306 - 325306, XP055218315 [Y] 10,13 * page 307 * DOI: http://dx.doi.org/10.1128/MMBR.69.2.306-325.2005 | [A] - S. A. F. T. VAN HIJUM ET AL, "Structure-Function Relationships of Glucansucrase and Fructansucrase Enzymes from Lactic Acid Bacteria", MICROBIOLOGY AND MOLECULAR BIOLOGY REVIEWS, (200603), vol. 70, no. 1, doi:10.1128/MMBR.70.1.157-176.2006, ISSN 1092-2172, pages 157 - 176, XP055179067 [A] 1-13 * the whole document * DOI: http://dx.doi.org/10.1128/MMBR.70.1.157-176.2006 | [XP] - "SubName: Full=Alternansucrase {ECO:0000313|EMBL:CDX66895.1}; EC=2.4.1.140 {ECO:0000313|EMBL:CDX66895.1};", UniProt, (20150204), Database accession no. A0A0A1IPZ8, URL: EBI, XP002745403 [XP] 1 * sequence . * | by applicant | US5229277 | EP2365084 | - MAGE; KABAT, "Immunochemical studies on dextrans", III. THE SPECIFICITIES OF RABBIT ANTIDEXTRANS, (1963), vol. 91, pages 634 - 640 | - LEMHUIS ET AL., "Glucansucrases: Three-dimensional structures, reactions, mechanism, alpha-glucan analysis and their implications in biotechnology and food applications", JOURNAL OF BIOTECHNOLOGY, (201301), vol. 163, no. 2, doi:doi:10.1016/j.jbiotec.2012.06.037, pages 250 - 272, XP055102852 DOI: http://dx.doi.org/10.1016/j.jbiotec.2012.06.037 | - SUMNER; HOWELL, "A method for determination of invertase activity", JOURNAL OF BIOLOGICAL CHEMISTRY, (1935), vol. 108, no. 51 | - "One-step synthesis of isomalto-oligosaccharide syrups and dextrans of controlled size using engineered dextransucrase", MOULIS C; VACA MEDINA G; SUWANNARANGSEE S; MONSAN P; REMAUD-SIMEON M; POTOCKI-VERONESE G, Biocatal. Biotransformation, TAYLOR & FRANCIS, (2008), pages 141 - 151 | - KIM ET AL., "Dextran molecular size and degree of branching as a function of sucrose concentration, pH, and temperature of reaction of Leuconostoc mesenteroides B-512FMCM dextransucrase", CARBOHYDRATE RESEARCH, (2003), vol. 338, doi:doi:10.1016/S0008-6215(03)00148-4, pages 1183 - 1189, XP004424302 DOI: http://dx.doi.org/10.1016/S0008-6215(03)00148-4 | - MOULIS ET AL., "High-level production and purification of a fully active recombinant dextransucrase from Leuconostoc mesenteroides NRRL B-512F", FEMS MICROBIOLOGY LETTERS, (200608), vol. 261, no. 2, doi:doi:10.1111/j.1574-6968.2006.00347.x, pages 203 - 210, XP002406211 DOI: http://dx.doi.org/10.1111/j.1574-6968.2006.00347.x | - KOTHARI ET AL., "Structural Characterization of Enzymatically Synthesized Dextran and Oligosaccharides from Leuconostoc mesenteroides NRRL B-1426 Dextransucrase. ISSN 0006-2979", BIOCHEMISTRY (MOSCOW, (2013), vol. 78, no. 10, pages 1164 - 1170 | - AMARI ET AL., "Characterization of a novel dextransucrase from Weissella confusa isolated from sourdough", APPL MICROBIOL BIOTECHNOL, (2013), vol. 97, pages 5413 - 5422 | - KRALJ ET AL., "Glucan synthesis in the genus Lactobacillus: isolation and characterization of glucansucrase genes, enzymes and glucan products from six different strains", MICROBIOLOGY, (2004), vol. 150, doi:doi:10.1099/mic.0.27321-0, pages 3681 - 3690, XP002360114 DOI: http://dx.doi.org/10.1099/mic.0.27321-0 | - STRIEGEL ET AL., "An SEC/MALS study of alternan degradation during size-exclusion chromatographic analysis", ANAL BIOANAL CHEM, (2009), vol. 394, doi:doi:10.1007/s00216-009-2895-5, pages 1887 - 1893, XP019736608 DOI: http://dx.doi.org/10.1007/s00216-009-2895-5 | - GOULAS, A. K. ET AL., "Synthesis of isomaltoligosaccharides and oligodextrans by the combined use of dextransucrase and dextransase", ENZYME MICROB. TECHNOL., (2004), vol. 35, pages 327 - 338 | - NAESSENS ET AL., "Leuconostoc dextransucrase and dextran: production, properties and applications", J. CHEM. TECHNOL. BIOTECHNOL., (2005), vol. 80, doi:doi:10.1002/jctb.1322, pages 845 - 860, XP001595980 DOI: http://dx.doi.org/10.1002/jctb.1322 | - EGGLESTON; COTE, G. L, "Oligosaccharides in food and agriculture", WASHINGTON, DC: ACS SYMPOSIUM SERIES 849, (2003), pages 1 - 15 | - THOMAS HEINZE; TIM LIEBERT; BRIGITTE HEUBLEIN; STEPHANIE HORNIG, "Functional Polymers Based on Dextran", ADV POLYM SCI, (2006), vol. 205, doi:doi:10.1007/12_100, pages 199 - 291, XP009138969 DOI: http://dx.doi.org/10.1007/12_100 |