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Extract from the Register of European Patents

EP About this file: EP3198005

EP3198005 - METHODS OF IN VITRO OOCYTE DEVELOPMENT [Right-click to bookmark this link]
StatusNo opposition filed within time limit
Status updated on  21.01.2022
Database last updated on 27.07.2024
FormerThe patent has been granted
Status updated on  12.02.2021
FormerGrant of patent is intended
Status updated on  11.10.2020
FormerExamination is in progress
Status updated on  09.11.2018
FormerRequest for examination was made
Status updated on  30.06.2017
FormerThe international publication has been made
Status updated on  14.04.2017
Most recent event   Tooltip27.04.2024Lapse of the patent in a contracting state
New state(s): MK		published on 29.05.2024  [2024/22]
Applicant(s)For all designated states
The University Court of the University of Edinburgh
Old College
South Bridge
Edinburgh
Midlothian EH8 9YL / GB
[2017/31]
Inventor(s)01 / TELFER, Evelyn Elizabeth
c/o The University Court of the University of
Edinburgh
Old College
South Bridge
Midlothian Central Scotland EH8 9YL / GB
02 / MCLAUGHLIN, Marie
c/o The University Court of the University of
Edinburgh
Old College
South Bridge
Midlothian Central Scotland EH8 9YL / GB
 [2021/11]
Former [2017/31]01 / TELFER, Evelyn Elizabeth
c/o The Univeristy Court of the Univiersity of
Edinburgh
Old College
South Bridge
Midlothian Central Scotland EH8 9YL / GB
02 / MCLAUGHLIN, Marie
c/o The University Court of the University of
Edinburgh
Old College
South Bridge
Midlothian Central Scotland EH8 9YL / GB
Representative(s)HGF
HGF Limited
1 City Walk
Leeds LS11 9DX / GB
[N/P]
Former [2021/11]HGF
1 City Walk
Leeds LS11 9DX / GB
Former [2017/31]HGF Limited
Delta House
50 West Nile Street
Glasgow G1 2NP / GB
Application number, filing date15777723.624.09.2015
[2017/31]
WO2015GB52769
Priority number, dateGB2014001685824.09.2014         Original published format: GB 201416858
[2017/31]
Filing languageEN
Procedural languageEN
PublicationType: A1 Application with search report
No.:WO2016046557
Date:31.03.2016
Language:EN
[2016/13]
Type: A1 Application with search report 
No.:EP3198005
Date:02.08.2017
Language:EN
The application published by WIPO in one of the EPO official languages on 31.03.2016 takes the place of the publication of the European patent application.
[2017/31]
Type: B1 Patent specification 
No.:EP3198005
Date:17.03.2021
Language:EN
[2021/11]
Search report(s)International search report - published on:EP31.03.2016
ClassificationIPC:C12N5/075, C12N5/071
[2017/31]
CPC:
C12N5/0609 (EP,US); C12N5/0611 (US); C12N5/0037 (US);
C12N5/0081 (US); C12N5/0682 (EP,US); A01N1/0268 (US);
C12N2500/25 (EP,US); C12N2500/38 (EP,US); C12N2500/90 (EP,US);
C12N2501/105 (EP,US); C12N2501/11 (EP,US); C12N2501/125 (EP,US);
C12N2501/16 (EP,US); C12N2501/31 (EP,US); C12N2501/392 (EP,US);
C12N2502/243 (EP,US); C12N2527/00 (EP,US); C12N5/0018 (US) (-)
Designated contracting statesAL,   AT,   BE,   BG,   CH,   CY,   CZ,   DE,   DK,   EE,   ES,   FI,   FR,   GB,   GR,   HR,   HU,   IE,   IS,   IT,   LI,   LT,   LU,   LV,   MC,   MK,   MT,   NL,   NO,   PL,   PT,   RO,   RS,   SE,   SI,   SK,   SM,   TR [2017/31]
Extension statesBANot yet paid
MENot yet paid
Validation statesMANot yet paid
TitleGerman:VERFAHREN ZUR IN-VITRO-OOZYTENENTWICKLUNG[2017/31]
English:METHODS OF IN VITRO OOCYTE DEVELOPMENT[2017/31]
French:PROCÉDÉS DE DÉVELOPPEMENT D'OVOCYTES IN VITRO[2017/31]
Entry into regional phase13.04.2017National basic fee paid 
13.04.2017Designation fee(s) paid 
13.04.2017Examination fee paid 
Examination procedure13.04.2017Examination requested  [2017/31]
13.04.2017Date on which the examining division has become responsible
14.11.2017Amendment by applicant (claims and/or description)
07.11.2018Despatch of a communication from the examining division (Time limit: M06)
30.04.2019Reply to a communication from the examining division
12.10.2020Communication of intention to grant the patent
05.02.2021Fee for grant paid
05.02.2021Fee for publishing/printing paid
05.02.2021Receipt of the translation of the claim(s)
Opposition(s)20.12.2021No opposition filed within time limit [2022/08]
Fees paidRenewal fee
05.09.2017Renewal fee patent year 03
24.09.2018Renewal fee patent year 04
29.07.2019Renewal fee patent year 05
14.09.2020Renewal fee patent year 06
Opt-out from the exclusive  Tooltip
competence of the Unified
Patent Court		 See the Register of the Unified Patent Court for opt-out data
Responsibility for the accuracy, completeness or quality of the data displayed under the link provided lies entirely with the Unified Patent Court.
Lapses during opposition  TooltipHU24.09.2015
AL17.03.2021
AT17.03.2021
CY17.03.2021
CZ17.03.2021
EE17.03.2021
ES17.03.2021
FI17.03.2021
HR17.03.2021
IT17.03.2021
LT17.03.2021
LV17.03.2021
MC17.03.2021
MK17.03.2021
PL17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SI17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
IS17.07.2021
PT19.07.2021
IE24.09.2021
LU24.09.2021
BE30.09.2021
CH30.09.2021
LI30.09.2021
[2024/22]
Former [2023/30]HU24.09.2015
AL17.03.2021
AT17.03.2021
CY17.03.2021
CZ17.03.2021
EE17.03.2021
ES17.03.2021
FI17.03.2021
HR17.03.2021
IT17.03.2021
LT17.03.2021
LV17.03.2021
MC17.03.2021
PL17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SI17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
IS17.07.2021
PT19.07.2021
IE24.09.2021
LU24.09.2021
BE30.09.2021
CH30.09.2021
LI30.09.2021
Former [2023/29]HU24.09.2015
AL17.03.2021
AT17.03.2021
CZ17.03.2021
EE17.03.2021
ES17.03.2021
FI17.03.2021
HR17.03.2021
IT17.03.2021
LT17.03.2021
LV17.03.2021
MC17.03.2021
PL17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SI17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
IS17.07.2021
PT19.07.2021
IE24.09.2021
LU24.09.2021
BE30.09.2021
CH30.09.2021
LI30.09.2021
Former [2022/37]AL17.03.2021
AT17.03.2021
CZ17.03.2021
EE17.03.2021
ES17.03.2021
FI17.03.2021
HR17.03.2021
IT17.03.2021
LT17.03.2021
LV17.03.2021
MC17.03.2021
PL17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SI17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
IS17.07.2021
PT19.07.2021
IE24.09.2021
LU24.09.2021
BE30.09.2021
CH30.09.2021
LI30.09.2021
Former [2022/35]AL17.03.2021
AT17.03.2021
CZ17.03.2021
EE17.03.2021
ES17.03.2021
FI17.03.2021
HR17.03.2021
IT17.03.2021
LT17.03.2021
LV17.03.2021
MC17.03.2021
PL17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SI17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
IS17.07.2021
PT19.07.2021
IE24.09.2021
LU24.09.2021
BE30.09.2021
Former [2022/34]AL17.03.2021
AT17.03.2021
CZ17.03.2021
EE17.03.2021
ES17.03.2021
FI17.03.2021
HR17.03.2021
IT17.03.2021
LT17.03.2021
LV17.03.2021
MC17.03.2021
PL17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SI17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
IS17.07.2021
PT19.07.2021
LU24.09.2021
BE30.09.2021
Former [2022/25]AL17.03.2021
AT17.03.2021
CZ17.03.2021
EE17.03.2021
ES17.03.2021
FI17.03.2021
HR17.03.2021
IT17.03.2021
LT17.03.2021
LV17.03.2021
MC17.03.2021
PL17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SI17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
IS17.07.2021
PT19.07.2021
Former [2022/23]AL17.03.2021
AT17.03.2021
CZ17.03.2021
EE17.03.2021
ES17.03.2021
FI17.03.2021
HR17.03.2021
IT17.03.2021
LT17.03.2021
LV17.03.2021
PL17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SI17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
IS17.07.2021
PT19.07.2021
Former [2022/13]AL17.03.2021
AT17.03.2021
CZ17.03.2021
EE17.03.2021
ES17.03.2021
FI17.03.2021
HR17.03.2021
LT17.03.2021
LV17.03.2021
PL17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SI17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
PT19.07.2021
Former [2022/10]AL17.03.2021
AT17.03.2021
CZ17.03.2021
EE17.03.2021
ES17.03.2021
FI17.03.2021
HR17.03.2021
LT17.03.2021
LV17.03.2021
PL17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
PT19.07.2021
Former [2022/08]AL17.03.2021
AT17.03.2021
CZ17.03.2021
EE17.03.2021
ES17.03.2021
FI17.03.2021
HR17.03.2021
LT17.03.2021
LV17.03.2021
PL17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
IS17.07.2021
PT19.07.2021
Former [2022/07]AL17.03.2021
AT17.03.2021
CZ17.03.2021
EE17.03.2021
FI17.03.2021
HR17.03.2021
LT17.03.2021
LV17.03.2021
PL17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
IS17.07.2021
PT19.07.2021
Former [2021/52]AT17.03.2021
CZ17.03.2021
EE17.03.2021
FI17.03.2021
HR17.03.2021
LT17.03.2021
LV17.03.2021
PL17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
IS17.07.2021
PT19.07.2021
Former [2021/51]AT17.03.2021
CZ17.03.2021
EE17.03.2021
FI17.03.2021
HR17.03.2021
LT17.03.2021
LV17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
IS17.07.2021
PT19.07.2021
Former [2021/50]AT17.03.2021
CZ17.03.2021
EE17.03.2021
FI17.03.2021
HR17.03.2021
LT17.03.2021
LV17.03.2021
RO17.03.2021
RS17.03.2021
SE17.03.2021
SK17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
PT19.07.2021
Former [2021/48]AT17.03.2021
CZ17.03.2021
EE17.03.2021
FI17.03.2021
HR17.03.2021
LT17.03.2021
LV17.03.2021
RS17.03.2021
SE17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
Former [2021/46]AT17.03.2021
FI17.03.2021
HR17.03.2021
LT17.03.2021
LV17.03.2021
RS17.03.2021
SE17.03.2021
SM17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
Former [2021/38]FI17.03.2021
HR17.03.2021
LV17.03.2021
RS17.03.2021
SE17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
Former [2021/37]FI17.03.2021
HR17.03.2021
LV17.03.2021
RS17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
Former [2021/36]FI17.03.2021
HR17.03.2021
BG17.06.2021
NO17.06.2021
GR18.06.2021
Former [2021/35]FI17.03.2021
HR17.03.2021
NO17.06.2021
Former [2021/33]FI17.03.2021
NO17.06.2021
Cited inInternational search[XYI]  - R. A. ANDERSON ET AL, "The immature human ovary shows loss of abnormal follicles and increasing follicle developmental competence through childhood and adolescence", HUMAN REPRODUCTION, GB, (20131017), vol. 29, no. 1, doi:10.1093/humrep/det388, ISSN 0268-1161, pages 97 - 106, XP055235522 [X] 21-23,25,42,43,45-47 * the whole document * [Y] 36-38 [I] 1-20,27-30

DOI:   http://dx.doi.org/10.1093/humrep/det388
 [XYI]  - M. MCLAUGHLIN ET AL, "Oocyte development in bovine primordial follicles is promoted by activin and FSH within a two-step serum-free culture system", REPRODUCTION, GB, (20100305), vol. 139, no. 6, doi:10.1530/REP-10-0025, ISSN 1470-1626, pages 971 - 978, XP055235388 [X] 21-25,31-34,48,49 * the whole document * [Y] 35 [I] 26

DOI:   http://dx.doi.org/10.1530/REP-10-0025
 [Y]  - Evelyn Telfer, "Novel systems and factors to improve oocyte quality in vitro", (20120329), URL: https://www.eshre.eu/~/media/sitecore-files/SIGs/Embryology/Archive/Stresa-2012/Telfer2.pdf?la=en, (20151211), XP055235634 [Y] 35-38 * the whole document *
 [X]  - Talal El-Hefnawy ET AL, "Synergism Between FSH and Activin in the Regulation of Proliferating Cell Nuclear Antigen (PCNA) and Cyclin D2 Expression in Rat Granulosa Cells", Endocrinology, doi:10.1210/endo.142.10.8438, (20011001), pages 4357 - 4362, URL: http://press.endocrine.org/doi/pdf/10.1210/endo.142.10.8438, (20151211), XP055235727 [X] 42-47 * the whole document *

DOI:   http://dx.doi.org/10.1210/endo.142.10.8438
 [X]  - SEUNG-HEE SONG ET AL, "Characterization of Porcine Multipotent Stem/Stromal Cells Derived from Skin, Adipose, and Ovarian Tissues and Their Differentiation In Vitro into Putative Oocyte-Like Cells", STEM CELLS AND DEVELOPMENT, NL, (20110801), vol. 20, no. 8, doi:10.1089/scd.2010.0203, ISSN 1547-3287, pages 1359 - 1370, XP055235890 [X] 39-41,50-52 * the whole document *

DOI:   http://dx.doi.org/10.1089/scd.2010.0203
 [T]  - EVELYN E. TELFER ET AL, "Strategies to support human oocyte development in vitro", THE INTERNATIONAL JOURNAL OF DEVELOPMENTAL BIOLOGY, (20120101), vol. 56, no. 10-11-12, doi:10.1387/ijdb.130001et, ISSN 0214-6282, pages 901 - 907, XP055092892 [T] 1-52 * the whole document *

DOI:   http://dx.doi.org/10.1387/ijdb.130001et
The EPO accepts no responsibility for the accuracy of data originating from other authorities; in particular, it does not guarantee that it is complete, up to date or fit for specific purposes.