blank Quick help
blank Maintenance news

Scheduled maintenance

Regular maintenance outages:
between 05.00 and 05.15 hrs CET (Monday to Sunday).

Other outages
Availability
Register Forum

2022.02.11

More...
blank News flashes

News flashes

New version of the European Patent Register - SPC information for Unitary Patents.

2024-03-06

More...
blank Related links

Extract from the Register of European Patents

EP About this file: EP2135503

EP2135503 - METHOD FOR IMPROVEMENT OF EFFICIENCY OF TRANSFORMATION IN PLANT, COMPRISING CO-CULTURE STEP FOR CULTURING PLANT TISSUE IN CO-CULTURE MEDIUM CONTAINING 3,6-DICHLORO-o-ANISIC ACID [Right-click to bookmark this link]
StatusNo opposition filed within time limit
Status updated on  28.11.2014
Database last updated on 15.07.2024
Most recent event   Tooltip15.07.2016Lapse of the patent in a contracting state
New state(s): LU, TR
published on 17.08.2016  [2016/33]
Applicant(s)For all designated states
Japan Tobacco, Inc.
2-1, Toranomon 2-chome, Minato-ku
Tokyo 105-8422 / JP
[2009/52]
Inventor(s)01 / ISHIDA, Yuji
c/o Japan Tobacco Inc.
Plant Innovation Center
700 Higashibara
Iwata-shi Shizuoka 438-0802 / JP
02 / HIEI, Yukoh
c/o Japan Tobacco Inc.
Plant Innovation Center
700 Higashibara
Iwata-shi Shizuoka 438-0802 / JP
 [2014/04]
Former [2009/52]01 / ISHIDA, Yuji
c/o Japan Tobacco Inc. Plant Innovation Center 700 Higashibara
Iwata-shi Shizuoka 438-0802 / JP
02 / HIEI, Yukoh
c/o Japan Tobacco Inc. Plant Innovation Center 700 Higashibara
Iwata-shi Shizuoka 438-0802 / JP
Representative(s)Isarpatent
Patent- und Rechtsanwälte Barth
Charles Hassa Peckmann & Partner mbB
Postfach 44 01 51
80750 München / DE
[N/P]
Former [2012/18]Isarpatent
Patent- und Rechtsanwälte
Postfach 44 01 51
80750 München / DE
Former [2009/52]Behnisch, Werner
Reinhard, Skuhra, Weise & Partner GbR Patent- und Rechtsanwälte Friedrichstrasse 31
80801 München / DE
Application number, filing date08721007.628.02.2008
[2009/52]
WO2008JP53558
Priority number, dateJP2007004916128.02.2007         Original published format: JP 2007049161
[2009/52]
Filing languageJA
Procedural languageEN
PublicationType: A1 Application with search report
No.:WO2008105508
Date:04.09.2008
Language:JA
[2008/36]
Type: A1 Application with search report 
No.:EP2135503
Date:23.12.2009
Language:EN
[2009/52]
Type: B1 Patent specification 
No.:EP2135503
Date:22.01.2014
Language:EN
[2014/04]
Search report(s)International search report - published on:JP04.09.2008
(Supplementary) European search report - dispatched on:EP30.06.2010
ClassificationIPC:A01H1/00, C12N15/09
[2009/52]
CPC:
C12N15/8205 (EP,US)
Designated contracting statesAT,   BE,   BG,   CH,   CY,   CZ,   DE,   DK,   EE,   ES,   FI,   FR,   GB,   GR,   HR,   HU,   IE,   IS,   IT,   LI,   LT,   LU,   LV,   MC,   MT,   NL,   NO,   PL,   PT,   RO,   SE,   SI,   SK,   TR [2009/52]
TitleGerman:VERFAHREN FÜR DIE VERBESSERUNG DER TRANSFORMATIONSEFFIZIENZ BEI PFLANZEN, UMFASSEND EINEN COKULTURSCHRITT FÜR DIE KULTIVIERUNG VON PFLANZENGEWEBE IN CO-KULTURMEDIUM ENTHALTEND 3,6-DICHLOR-O-ANISSÄURE[2009/52]
English:METHOD FOR IMPROVEMENT OF EFFICIENCY OF TRANSFORMATION IN PLANT, COMPRISING CO-CULTURE STEP FOR CULTURING PLANT TISSUE IN CO-CULTURE MEDIUM CONTAINING 3,6-DICHLORO-o-ANISIC ACID[2009/52]
French:PROCÉDÉ D'AMÉLIORATION DE L'EFFICACITÉ DE LA TRANSFORMATION DANS UNE PLANTE, COMPRENANT UNE ÉTAPE DE CO-CULTURE POUR METTRE EN CULTURE UN TISSU VÉGÉTAL DANS UN MILIEU DE CO-CULTURE COMPRENANT DE L'ACIDE 3,6-DICHLORO-o-ANISIQUE[2009/52]
Entry into regional phase23.09.2009Translation filed 
23.09.2009National basic fee paid 
23.09.2009Search fee paid 
23.09.2009Designation fee(s) paid 
23.09.2009Examination fee paid 
Examination procedure23.09.2009Examination requested  [2009/52]
31.01.2011Amendment by applicant (claims and/or description)
25.08.2011Despatch of a communication from the examining division (Time limit: M06)
05.03.2012Reply to a communication from the examining division
18.06.2012Despatch of a communication from the examining division (Time limit: M06)
18.12.2012Reply to a communication from the examining division
23.09.2013Communication of intention to grant the patent
10.12.2013Fee for grant paid
10.12.2013Fee for publishing/printing paid
Divisional application(s)The date of the Examining Division's first communication in respect of the earliest application for which a communication has been issued is  25.08.2011
Opposition(s)23.10.2014No opposition filed within time limit [2015/01]
Fees paidRenewal fee
26.02.2010Renewal fee patent year 03
28.02.2011Renewal fee patent year 04
29.02.2012Renewal fee patent year 05
28.02.2013Renewal fee patent year 06
Opt-out from the exclusive  Tooltip
competence of the Unified
Patent Court
See the Register of the Unified Patent Court for opt-out data
Responsibility for the accuracy, completeness or quality of the data displayed under the link provided lies entirely with the Unified Patent Court.
Lapses during opposition  TooltipHU28.02.2008
AT22.01.2014
BG22.01.2014
CY22.01.2014
CZ22.01.2014
DK22.01.2014
EE22.01.2014
ES22.01.2014
FI22.01.2014
HR22.01.2014
IT22.01.2014
LT22.01.2014
LV22.01.2014
MC22.01.2014
MT22.01.2014
NL22.01.2014
PL22.01.2014
RO22.01.2014
SE22.01.2014
SI22.01.2014
SK22.01.2014
TR22.01.2014
CH28.02.2014
IE28.02.2014
LI28.02.2014
LU28.02.2014
GB22.04.2014
NO22.04.2014
GR23.04.2014
IS22.05.2014
PT22.05.2014
[2016/33]
Former [2016/32]HU28.02.2008
AT22.01.2014
BG22.01.2014
CY22.01.2014
CZ22.01.2014
DK22.01.2014
EE22.01.2014
ES22.01.2014
FI22.01.2014
HR22.01.2014
IT22.01.2014
LT22.01.2014
LV22.01.2014
MC22.01.2014
MT22.01.2014
NL22.01.2014
PL22.01.2014
RO22.01.2014
SE22.01.2014
SI22.01.2014
SK22.01.2014
CH28.02.2014
IE28.02.2014
LI28.02.2014
GB22.04.2014
NO22.04.2014
GR23.04.2014
IS22.05.2014
PT22.05.2014
Former [2016/29]AT22.01.2014
BG22.01.2014
CY22.01.2014
CZ22.01.2014
DK22.01.2014
EE22.01.2014
ES22.01.2014
FI22.01.2014
HR22.01.2014
IT22.01.2014
LT22.01.2014
LV22.01.2014
MC22.01.2014
MT22.01.2014
NL22.01.2014
PL22.01.2014
RO22.01.2014
SE22.01.2014
SI22.01.2014
SK22.01.2014
CH28.02.2014
IE28.02.2014
LI28.02.2014
GB22.04.2014
NO22.04.2014
GR23.04.2014
IS22.05.2014
PT22.05.2014
Former [2016/28]AT22.01.2014
BG22.01.2014
CY22.01.2014
CZ22.01.2014
DK22.01.2014
EE22.01.2014
ES22.01.2014
FI22.01.2014
HR22.01.2014
LT22.01.2014
LV22.01.2014
MC22.01.2014
MT22.01.2014
NL22.01.2014
PL22.01.2014
RO22.01.2014
SE22.01.2014
SI22.01.2014
SK22.01.2014
CH28.02.2014
IE28.02.2014
LI28.02.2014
GB22.04.2014
NO22.04.2014
GR23.04.2014
IS22.05.2014
PT22.05.2014
Former [2016/25]AT22.01.2014
BG22.01.2014
CY22.01.2014
CZ22.01.2014
DK22.01.2014
EE22.01.2014
ES22.01.2014
FI22.01.2014
HR22.01.2014
LT22.01.2014
LV22.01.2014
MC22.01.2014
MT22.01.2014
NL22.01.2014
PL22.01.2014
RO22.01.2014
SE22.01.2014
SI22.01.2014
SK22.01.2014
CH28.02.2014
IE28.02.2014
LI28.02.2014
GB22.04.2014
NO22.04.2014
IS22.05.2014
PT22.05.2014
Former [2016/12]AT22.01.2014
CY22.01.2014
CZ22.01.2014
DK22.01.2014
EE22.01.2014
ES22.01.2014
FI22.01.2014
HR22.01.2014
LT22.01.2014
LV22.01.2014
MC22.01.2014
MT22.01.2014
NL22.01.2014
PL22.01.2014
RO22.01.2014
SE22.01.2014
SI22.01.2014
SK22.01.2014
CH28.02.2014
IE28.02.2014
LI28.02.2014
GB22.04.2014
NO22.04.2014
IS22.05.2014
PT22.05.2014
Former [2015/24]AT22.01.2014
CY22.01.2014
CZ22.01.2014
DK22.01.2014
EE22.01.2014
ES22.01.2014
FI22.01.2014
HR22.01.2014
LT22.01.2014
LV22.01.2014
MC22.01.2014
NL22.01.2014
PL22.01.2014
RO22.01.2014
SE22.01.2014
SI22.01.2014
SK22.01.2014
CH28.02.2014
IE28.02.2014
LI28.02.2014
GB22.04.2014
NO22.04.2014
IS22.05.2014
PT22.05.2014
Former [2015/09]AT22.01.2014
CY22.01.2014
CZ22.01.2014
DK22.01.2014
EE22.01.2014
ES22.01.2014
FI22.01.2014
HR22.01.2014
LT22.01.2014
LV22.01.2014
MC22.01.2014
NL22.01.2014
PL22.01.2014
RO22.01.2014
SE22.01.2014
SK22.01.2014
CH28.02.2014
IE28.02.2014
LI28.02.2014
GB22.04.2014
NO22.04.2014
IS22.05.2014
PT22.05.2014
Former [2015/07]AT22.01.2014
CY22.01.2014
CZ22.01.2014
DK22.01.2014
EE22.01.2014
ES22.01.2014
FI22.01.2014
HR22.01.2014
LT22.01.2014
LV22.01.2014
MC22.01.2014
NL22.01.2014
PL22.01.2014
RO22.01.2014
SE22.01.2014
SK22.01.2014
CH28.02.2014
IE28.02.2014
LI28.02.2014
NO22.04.2014
IS22.05.2014
PT22.05.2014
Former [2014/50]AT22.01.2014
CY22.01.2014
CZ22.01.2014
DK22.01.2014
EE22.01.2014
ES22.01.2014
FI22.01.2014
HR22.01.2014
LT22.01.2014
LV22.01.2014
MC22.01.2014
NL22.01.2014
PL22.01.2014
RO22.01.2014
SE22.01.2014
SK22.01.2014
CH28.02.2014
LI28.02.2014
NO22.04.2014
IS22.05.2014
PT22.05.2014
Former [2014/49]AT22.01.2014
CY22.01.2014
CZ22.01.2014
DK22.01.2014
EE22.01.2014
ES22.01.2014
FI22.01.2014
HR22.01.2014
LT22.01.2014
LV22.01.2014
MC22.01.2014
NL22.01.2014
RO22.01.2014
SE22.01.2014
CH28.02.2014
LI28.02.2014
NO22.04.2014
IS22.05.2014
PT22.05.2014
Former [2014/48]AT22.01.2014
CY22.01.2014
CZ22.01.2014
DK22.01.2014
EE22.01.2014
ES22.01.2014
FI22.01.2014
HR22.01.2014
LT22.01.2014
LV22.01.2014
MC22.01.2014
NL22.01.2014
SE22.01.2014
CH28.02.2014
LI28.02.2014
NO22.04.2014
IS22.05.2014
PT22.05.2014
Former [2014/46]AT22.01.2014
CY22.01.2014
ES22.01.2014
FI22.01.2014
HR22.01.2014
LT22.01.2014
LV22.01.2014
MC22.01.2014
NL22.01.2014
SE22.01.2014
CH28.02.2014
LI28.02.2014
NO22.04.2014
IS22.05.2014
PT22.05.2014
Former [2014/41]AT22.01.2014
CY22.01.2014
ES22.01.2014
FI22.01.2014
HR22.01.2014
LT22.01.2014
LV22.01.2014
NL22.01.2014
SE22.01.2014
NO22.04.2014
IS22.05.2014
PT22.05.2014
Former [2014/39]AT22.01.2014
CY22.01.2014
ES22.01.2014
FI22.01.2014
LT22.01.2014
NL22.01.2014
SE22.01.2014
NO22.04.2014
IS22.05.2014
PT22.05.2014
Former [2014/37]AT22.01.2014
CY22.01.2014
ES22.01.2014
FI22.01.2014
LT22.01.2014
SE22.01.2014
NO22.04.2014
IS22.05.2014
PT22.05.2014
Former [2014/36]LT22.01.2014
NO22.04.2014
IS22.05.2014
Documents cited:Search[X]JP2000342253  (JAPAN TOBACCO INC) [X] 1-8* paragraph [0042] - paragraph [0055] *
International search[Y]JP2000342253  (JAPAN TOBACCO INC);
 [X]  - FRAME B.R. ET AL., "Improved Agrobacterium-mediated transformation of three maize inbred lines using MS salts", PLANT CELL REP., (2006), vol. 25, no. 10, pages 1024 - 1034, XP019423768

DOI:   http://dx.doi.org/10.1007/s00299-006-0145-2
 [X]  - TRIFONOVA S. ET AL., "Agrobacterium-mediated transgene delivery and integration into barley under a range of in vitro culture conditions", PLANT SCIENCE, (2001), vol. 161, no. 5, pages 871 - 880, XP002958562

DOI:   http://dx.doi.org/10.1016/S0168-9452(01)00479-4
 [A]  - HIEI Y. ET AL., "Improved protocols for transformation of indica rice mediated by Agrobacterium tumefaciens", PLANT CELL, TISSUE AND ORGAN CULTURE, (2006), vol. 85, no. 3, pages 271 - 283, XP019409331

DOI:   http://dx.doi.org/10.1007/s11240-005-9069-8
by applicantJP2000023675
 JP2000342255
 JP2000342256
 JP2000342253
 WO2005017169
 WO2005017152
 JP2007049161
    - DE CLEENE, M.; DE LEY, J., "The host range of crown gall", BOT. REV., (1976), vol. 42, pages 389 - 466
    - GRIMSLEY, N. ET AL., "Agrobacterium-mediated delivery of infectious maize streak virus into maize plants", NATURE, (1987), vol. 325, doi:doi:10.1038/325177a0, pages 177 - 179, XP002982470

DOI:   http://dx.doi.org/10.1038/325177a0
    - GRIMSLEY, N.H. ET AL., "Meristematic tissues of maize plants are most susceptible to Agroinfection with maize streak virus", BIO/TECHNOLOGY, (1988), vol. 6, pages 185 - 189
    - GRIMSLEY, N. ET AL., "DNA transfer from Agrobacterium to Zea mays or Brassica by agroinfection is dependent on bacterial virulence functions", MOL. GEN. GENET., (1989), vol. 217, pages 309 - 316
    - GOULD, J. ET AL., "Transformation of Zea mays L. using Agrobacterium tumefaciens and shoot apex", PLANT PHYSIOL., (1991), vol. 95, pages 426 - 434
    - MOONEY, P.A. ET AL., "Agrobacterium tumefaciens-gene transfer into wheat tissues", PLANT CELL, TISSUES AND ORGAN CULTURE, (1991), vol. 25, pages 209 - 218, XP000560763
    - RAINERI, D.M. ET AL., "Agrobacterium-mediated transformation of rice (Oryza sativa L.)", BIO/TECHNOLOGY, (1990), vol. 8, doi:doi:10.1038/nbt0190-33, pages 33 - 38, XP000918272

DOI:   http://dx.doi.org/10.1038/nbt0190-33
    - POTRYCUS, I, "Gene transfer to cereals: an assessment", BIO/TECHNOLOGY, (1990), vol. 8, pages 535 - 542
    - CHAN, M-T. ET AL., "Agrobacterium-mediated production of transgenic rice plants expressing a chimeric a-amylase promoter / (3-glucuronidase gene", PLANT MOL. BIOL., (1993), vol. 22, doi:doi:10.1007/BF00015978, pages 491 - 506, XP009006601

DOI:   http://dx.doi.org/10.1007/BF00015978
    - HIEI, Y. ET AL., "Efficient transformation of rice (Oryza sativa L.) mediated by Agrobacterium and sequence analysis of the boundaries of the T-DNA", THE PLANT JOURNAL, (1994), vol. 6, doi:doi:10.1046/j.1365-313X.1994.6020271.x, pages 271 - 282, XP002922710

DOI:   http://dx.doi.org/10.1046/j.1365-313X.1994.6020271.x
    - ISHIDA, Y. ET AL., "High efficiency transformation of maize (Zea mays L.) mediated by Agrobacterium tumefaciens", NATURE BIOTECHNOLOGY, (1996), vol. 14, doi:doi:10.1038/nbt0696-745, pages 745 - 750, XP002046364

DOI:   http://dx.doi.org/10.1038/nbt0696-745
    - CHENG, M. ET AL., "Genetic transformation of wheat mediated by Agrobacterium tumefaciens", PLANT PHYSIOL., (1997), vol. 115, pages 971 - 980, XP002918248
    - TINGAY, S. ET AL., "Agrobacterium tumefaciens-mediated barley transformation", PLANT J., (1997), vol. 11, doi:doi:10.1046/j.1365-313X.1997.11061369.x, pages 1369 - 1376, XP009080096

DOI:   http://dx.doi.org/10.1046/j.1365-313X.1997.11061369.x
    - ZHAO, Z.-Y. ET AL., "Agrobacterium-mediated sorghum transformation", PLANT MOL. BIOL., (2000), vol. 44, doi:doi:10.1023/A:1026507517182, pages 789 - 798, XP002533153

DOI:   http://dx.doi.org/10.1023/A:1026507517182
    - DEJI, A. ET AL., "Genomic organization and transcriptional regulation of maize ZmRRl and ZmRR2 encoding cytokinin-inducible response regulators", BIOCHIM. ET BIOPHYS. ACTA, (2000), vol. 1492, doi:doi:10.1016/S0167-4781(00)00076-2, pages 216 - 220, XP004275678

DOI:   http://dx.doi.org/10.1016/S0167-4781(00)00076-2
    - NEGROTTO, D. ET AL., "The use of phosphomannose-isomerase as a selection marker to recover transgenic maize plants (Zea mays L.) via Agrobacterium transformation", PLANT CELL REPORTS, (2000), vol. 19, pages 798 - 803
    - NOMURA, M. ET AL., "The evolution ofC4 plants: acquisition of cis-regulatory sequences in the promoter of C4-type pyruvate, orthophosphate dikinase gene", PLANT J., (2000), vol. 22, pages 211 - 221
    - NOMURA, M. ET AL., "The promoter ofrbcS in a C3 plant (rice) directs organ-specific, light-dependent expression in a C4 plant (maize), but does not confer bundle sheath cell-specific expression", PLANT MOL. BIOL., (2000), vol. 44, doi:doi:10.1023/A:1006461812053, pages 99 - 106, XP002179458

DOI:   http://dx.doi.org/10.1023/A:1006461812053
    - TANIGUCHI, M. ET AL., "The promoter for the maize C4 pyruvate, orthophosphate dikinase gene directs cell- and tissue-specific transcription in transgenic maize plants", PLANT CELL PHYSIOL., (2000), vol. 41, pages 42 - 48
    - ZHAO, Z.-Y. ET AL., "High throughput genetic transformation mediated by Agrobacterium tumefaciens in maize", MOL. BREED., (2001), vol. 8, doi:doi:10.1023/A:1015243600325, pages 323 - 333, XP001100034

DOI:   http://dx.doi.org/10.1023/A:1015243600325
    - FRAME, B.R. ET AL., "Agrobacterium tumefaciens-mediated transformation of maize embryos using a standard binary vector system", PLANT PHYSIOL., (2002), vol. 129, doi:doi:10.1104/pp.000653, pages 13 - 22, XP002263971

DOI:   http://dx.doi.org/10.1104/pp.000653
    - ISHIDA, Y. ET AL., "Improved protocol for transformation of maize (Zea mays L.) mediated by Agrobacterium tumefaciens", PLANT BIOTECHNOLOGY, (2003), vol. 20, pages 57 - 66, XP009070864
    - DUNCAN, D.R. ET AL., "The production of callus capable of plant regeneration from immature embryos of numerous Zea mays genotypes", PLANTA, (1985), vol. 165, doi:doi:10.1007/BF00392228, pages 322 - 332, XP000648041

DOI:   http://dx.doi.org/10.1007/BF00392228
    - ZHANG, W. ET AL., "Cre/lox-mediated marker gene excision in transgenic maize (Zea mays L.) plants", THEOR. APPL. GENET., (2003), vol. 107, pages 1157 - 1168
    - HUANG, X.; WEI, Z., "Successful Agrobacterium-mediated genetic transformation of maize elite inbred lines", PLANT CELL, TISSUE AND ORGAN CULTURE, (2005), vol. 83, pages 187 - 200, XP019268654
    - FRAME, B.R. ET AL., "Improved Agrobacterium-mediated transformation of three maize inbred lines using MS salts", PLANT CELL REP., (2006), vol. 25, doi:doi:10.1007/s00299-006-0145-2, pages 1024 - 1034, XP019423768

DOI:   http://dx.doi.org/10.1007/s00299-006-0145-2
    - KOMARI, T. ET AL., "Vectors carrying two separate T-DNAs for co-transformation of higher plants mediated by Agrobacterium tumefaciens and segregation of transformants free from selection markers", THE PLANT JOURNAL, (1996), vol. 10, doi:doi:10.1046/j.1365-313X.1996.10010165.x, pages 165 - 174, XP003024409

DOI:   http://dx.doi.org/10.1046/j.1365-313X.1996.10010165.x
    - KOMARI, T. ET AL., "Efficient selection of somatic hybrids in Nicotiana tabacum L. using a combination of drug-resistance merkers introduced by transformation", THEOR. APPL. GENET., (1989), vol. 77, pages 547 - 552
    - LINSMAIER, E.; SKOOG, F., "Organic growth factor requirements of tobacco tissue culture", PHYSIOL. PLANT., (1965), vol. 18, pages 100 - 127, XP000617350
    - "The N6 medium and its applications to anther culture of cereal crops", CHU, C.-C., Proc. Symp. Plant Tissue Culture, PEKING: SCIENCE PRESS, (1978), pages 43 - 50
    - WATSON, B. ET AL., "Plasmid required for virulence of Agrobacterium tumefaciens", J BACTERIOL, (1975), vol. 123, pages 255 - 264
The EPO accepts no responsibility for the accuracy of data originating from other authorities; in particular, it does not guarantee that it is complete, up to date or fit for specific purposes.