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Extract from the Register of European Patents

EP About this file: EP2420561

EP2420561 - ASPERGILLUS SP. HAVING LARGE-SCALE GENOME DUPLICATION [Right-click to bookmark this link]
StatusNo opposition filed within time limit
Status updated on  10.07.2020
Database last updated on 03.09.2024
FormerThe patent has been granted
Status updated on  26.07.2019
FormerGrant of patent is intended
Status updated on  02.04.2019
FormerExamination is in progress
Status updated on  16.05.2017
Most recent event   Tooltip08.07.2022Lapse of the patent in a contracting state
New state(s): MK		published on 10.08.2022  [2022/32]
Applicant(s)For all designated states
Kikkoman Corporation
250, Noda Noda-shi
Chiba 278-8601 / JP
[2012/08]
Inventor(s)01 / SATO, Atsushi
c/o Kikkoman Corporation
250, Noda
Noda-shi Chiba 278-8601 / JP
02 / HANYA, Yoshiki
c/o Kikkoman Corporation
250, Noda
Noda-shi Chiba 278-8601 / JP
 [2012/08]
Representative(s)Kramer Barske Schmidtchen Patentanwälte PartG mbB
European Patent Attorneys
Landsberger Strasse 300
80687 München / DE
[2019/35]
Former [2012/08]Kramer - Barske - Schmidtchen
Landsberger Strasse 300
80687 München / DE
Application number, filing date10764549.116.04.2010
[2019/35]
WO2010JP56880
Priority number, dateJP2009010064517.04.2009         Original published format: JP 2009100645
[2012/08]
Filing languageJA
Procedural languageEN
PublicationType: A1 Application with search report
No.:WO2010119967
Date:21.10.2010
Language:JA
[2010/42]
Type: A1 Application with search report 
No.:EP2420561
Date:22.02.2012
Language:EN
[2012/08]
Type: B1 Patent specification 
No.:EP2420561
Date:28.08.2019
Language:EN
[2019/35]
Search report(s)International search report - published on:JP21.10.2010
(Supplementary) European search report - dispatched on:EP02.01.2014
ClassificationIPC:C12R1/69, C12N9/62, C12R1/66, C12N9/30, A23L2/84, C12N9/58, C12N15/52, A23L27/50
[2019/14]
CPC:
A23L27/50 (EP,US); C12N1/145 (EP,US); C12N9/242 (EP,US);
C12N9/62 (EP,US); C12Y302/01001 (EP,US); A23L2/84 (US);
C12N15/52 (EP,US); C12N9/58 (EP,US); C12R2001/66 (EP,US);
C12R2001/69 (EP,US) (-)
Former IPC [2014/03]A23L1/238, C12R1/69, C12R1/66, C12N9/30, C12N9/62
Former IPC [2012/08]C12N1/14, A23L1/238, C12N15/01
Designated contracting statesAT,   BE,   BG,   CH,   CY,   CZ,   DE,   DK,   EE,   ES,   FI,   FR,   GB,   GR,   HR,   HU,   IE,   IS,   IT,   LI,   LT,   LU,   LV,   MC,   MK,   MT,   NL,   NO,   PL,   PT,   RO,   SE,   SI,   SK,   SM,   TR [2012/08]
Extension statesALNot yet paid
BANot yet paid
MENot yet paid
RSNot yet paid
TitleGerman:ASPERGILLUS SP. MIT GENOMDUPLIZIERUNG in großem Maßstab[2019/14]
English:ASPERGILLUS SP. HAVING LARGE-SCALE GENOME DUPLICATION[2012/08]
French:CELLULES DE LEVURE À DUPLICATION GÉNOMIQUE DE GRANDE ENVERGURE[2012/08]
Former [2012/08]ASPERGILLUS SP. MIT MASSENGENOMDUPLIZIERUNG
Biological materialThis application mentions deposited biological material, check the file for details
Entry into regional phase25.03.2011Translation filed 
25.03.2011National basic fee paid 
25.03.2011Search fee paid 
25.03.2011Designation fee(s) paid 
25.03.2011Examination fee paid 
Examination procedure25.03.2011Examination requested  [2012/08]
03.07.2014Amendment by applicant (claims and/or description)
17.02.2016Despatch of a communication from the examining division (Time limit: M04)
30.05.2016Reply to a communication from the examining division
08.05.2017Despatch of a communication from the examining division (Time limit: M04)
14.08.2017Reply to a communication from the examining division
03.04.2019Communication of intention to grant the patent
16.07.2019Fee for grant paid
16.07.2019Fee for publishing/printing paid
16.07.2019Receipt of the translation of the claim(s)
Divisional application(s)The date of the Examining Division's first communication in respect of the earliest application for which a communication has been issued is  17.02.2016
Opposition(s)03.06.2020No opposition filed within time limit [2020/33]
Fees paidRenewal fee
25.04.2012Renewal fee patent year 03
25.04.2013Renewal fee patent year 04
27.03.2014Renewal fee patent year 05
27.04.2015Renewal fee patent year 06
03.03.2016Renewal fee patent year 07
26.04.2017Renewal fee patent year 08
19.04.2018Renewal fee patent year 09
12.04.2019Renewal fee patent year 10
Opt-out from the exclusive  Tooltip
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Responsibility for the accuracy, completeness or quality of the data displayed under the link provided lies entirely with the Unified Patent Court.
Lapses during opposition  TooltipHU16.04.2010
AT28.08.2019
CY28.08.2019
CZ28.08.2019
DK28.08.2019
EE28.08.2019
ES28.08.2019
FI28.08.2019
HR28.08.2019
IT28.08.2019
LT28.08.2019
LV28.08.2019
MC28.08.2019
MK28.08.2019
MT28.08.2019
PL28.08.2019
RO28.08.2019
SE28.08.2019
SI28.08.2019
SK28.08.2019
SM28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
IS28.12.2019
PT30.12.2019
GB16.04.2020
IE16.04.2020
LU16.04.2020
BE30.04.2020
CH30.04.2020
FR30.04.2020
LI30.04.2020
[2022/32]
Former [2022/27]HU16.04.2010
AT28.08.2019
CY28.08.2019
CZ28.08.2019
DK28.08.2019
EE28.08.2019
ES28.08.2019
FI28.08.2019
HR28.08.2019
IT28.08.2019
LT28.08.2019
LV28.08.2019
MC28.08.2019
MT28.08.2019
PL28.08.2019
RO28.08.2019
SE28.08.2019
SI28.08.2019
SK28.08.2019
SM28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
IS28.12.2019
PT30.12.2019
GB16.04.2020
IE16.04.2020
LU16.04.2020
BE30.04.2020
CH30.04.2020
FR30.04.2020
LI30.04.2020
Former [2021/22]AT28.08.2019
CZ28.08.2019
DK28.08.2019
EE28.08.2019
ES28.08.2019
FI28.08.2019
HR28.08.2019
IT28.08.2019
LT28.08.2019
LV28.08.2019
MC28.08.2019
PL28.08.2019
RO28.08.2019
SE28.08.2019
SI28.08.2019
SK28.08.2019
SM28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
IS28.12.2019
PT30.12.2019
GB16.04.2020
IE16.04.2020
LU16.04.2020
BE30.04.2020
CH30.04.2020
FR30.04.2020
LI30.04.2020
Former [2021/20]AT28.08.2019
CZ28.08.2019
DK28.08.2019
EE28.08.2019
ES28.08.2019
FI28.08.2019
HR28.08.2019
IT28.08.2019
LT28.08.2019
LV28.08.2019
MC28.08.2019
PL28.08.2019
RO28.08.2019
SE28.08.2019
SI28.08.2019
SK28.08.2019
SM28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
IS28.12.2019
PT30.12.2019
IE16.04.2020
LU16.04.2020
BE30.04.2020
CH30.04.2020
FR30.04.2020
LI30.04.2020
Former [2021/12]AT28.08.2019
CZ28.08.2019
DK28.08.2019
EE28.08.2019
ES28.08.2019
FI28.08.2019
HR28.08.2019
IT28.08.2019
LT28.08.2019
LV28.08.2019
MC28.08.2019
PL28.08.2019
RO28.08.2019
SE28.08.2019
SI28.08.2019
SK28.08.2019
SM28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
IS28.12.2019
PT30.12.2019
LU16.04.2020
BE30.04.2020
CH30.04.2020
FR30.04.2020
LI30.04.2020
Former [2021/08]AT28.08.2019
CZ28.08.2019
DK28.08.2019
EE28.08.2019
ES28.08.2019
FI28.08.2019
HR28.08.2019
IT28.08.2019
LT28.08.2019
LV28.08.2019
MC28.08.2019
PL28.08.2019
RO28.08.2019
SE28.08.2019
SI28.08.2019
SK28.08.2019
SM28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
IS28.12.2019
PT30.12.2019
LU16.04.2020
CH30.04.2020
LI30.04.2020
Former [2020/52]AT28.08.2019
CZ28.08.2019
DK28.08.2019
EE28.08.2019
ES28.08.2019
FI28.08.2019
HR28.08.2019
IT28.08.2019
LT28.08.2019
LV28.08.2019
MC28.08.2019
PL28.08.2019
RO28.08.2019
SE28.08.2019
SI28.08.2019
SK28.08.2019
SM28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
IS28.12.2019
PT30.12.2019
Former [2020/37]AT28.08.2019
CZ28.08.2019
DK28.08.2019
EE28.08.2019
ES28.08.2019
FI28.08.2019
HR28.08.2019
IT28.08.2019
LT28.08.2019
LV28.08.2019
PL28.08.2019
RO28.08.2019
SE28.08.2019
SI28.08.2019
SK28.08.2019
SM28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
IS28.12.2019
PT30.12.2019
Former [2020/36]AT28.08.2019
CZ28.08.2019
DK28.08.2019
EE28.08.2019
ES28.08.2019
FI28.08.2019
HR28.08.2019
IT28.08.2019
LT28.08.2019
LV28.08.2019
PL28.08.2019
RO28.08.2019
SE28.08.2019
SK28.08.2019
SM28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
IS28.12.2019
PT30.12.2019
Former [2020/27]AT28.08.2019
CZ28.08.2019
DK28.08.2019
EE28.08.2019
ES28.08.2019
FI28.08.2019
HR28.08.2019
IT28.08.2019
LT28.08.2019
LV28.08.2019
PL28.08.2019
RO28.08.2019
SE28.08.2019
SK28.08.2019
SM28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
PT30.12.2019
IS24.02.2020
Former [2020/25]AT28.08.2019
CZ28.08.2019
DK28.08.2019
EE28.08.2019
ES28.08.2019
FI28.08.2019
HR28.08.2019
IT28.08.2019
LT28.08.2019
LV28.08.2019
PL28.08.2019
RO28.08.2019
SE28.08.2019
SK28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
PT30.12.2019
IS24.02.2020
Former [2020/24]AT28.08.2019
DK28.08.2019
EE28.08.2019
ES28.08.2019
FI28.08.2019
HR28.08.2019
IT28.08.2019
LT28.08.2019
LV28.08.2019
PL28.08.2019
RO28.08.2019
SE28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
PT30.12.2019
IS24.02.2020
Former [2020/23]AT28.08.2019
DK28.08.2019
EE28.08.2019
ES28.08.2019
FI28.08.2019
HR28.08.2019
IT28.08.2019
LT28.08.2019
LV28.08.2019
PL28.08.2019
RO28.08.2019
SE28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
IS28.12.2019
PT30.12.2019
Former [2020/22]DK28.08.2019
EE28.08.2019
ES28.08.2019
FI28.08.2019
HR28.08.2019
LT28.08.2019
LV28.08.2019
SE28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
IS28.12.2019
PT30.12.2019
Former [2020/17]ES28.08.2019
FI28.08.2019
HR28.08.2019
LT28.08.2019
LV28.08.2019
SE28.08.2019
TR28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
IS28.12.2019
PT30.12.2019
Former [2020/12]ES28.08.2019
FI28.08.2019
HR28.08.2019
LT28.08.2019
LV28.08.2019
SE28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
IS28.12.2019
PT30.12.2019
Former [2020/11]ES28.08.2019
FI28.08.2019
HR28.08.2019
LT28.08.2019
LV28.08.2019
SE28.08.2019
BG28.11.2019
NO28.11.2019
GR29.11.2019
PT30.12.2019
Former [2020/10]FI28.08.2019
LT28.08.2019
SE28.08.2019
BG28.11.2019
NO28.11.2019
PT30.12.2019
Former [2020/09]FI28.08.2019
LT28.08.2019
NO28.11.2019
Former [2020/08]LT28.08.2019
NO28.11.2019
Documents cited:Search[X]JPS57174087  ;
 [X]JPS60188057  ;
 [XPY]JP2009095279  ;
 [Y]US6830905  (KOIBUCHI KYOKO [JP], et al) [Y] 1-13 * column 8, line 64 - column 9, line 7 * * column 20, line 56 - line 59 *;
 [Y]US6090607  (VAN DEN BROEK PETER [CH], et al) [Y] 1-13 * column 4, line 40 - line 61 * * column 7, line 35 - line 45; examples 1, 3, 4, 5 *;
 [YP]JP2010075131
 [X]  - DATABASE WPI, 0, Derwent World Patents Index, vol. 1982, no. 48, Database accession no. 1982-03726J, XP002717406 & JPS57174087 A 19821026 (HIGETA SHOYU KK) [X] 1-5,8,9,11 * abstract *
 [XY]  - YOKOYAMA T ET AL, "Breeding koji mould by a combination of mutation and diploidization. (translated)", FSTA, INTERNATIONAL FOOD INFORMATION SERVICE (IFIS), FRANkFURT-MAIN, DE, Database accession no. FS-1984-04-T-0223, XP002717407 [X] 1-5,8,11 * abstract * [Y] 1-13
    [ ] - YOKOAMA T, KADOWAKI K, JOURNAL OF THE JAPAN SOY SAUCE RESEARCH INSTITUTE [NIHON SHOYU KENKUJO ZASSHI] 1983, (1983), vol. 9, no. 4, page 137, XP009174806
 [X]  - DATABASE WPI, 0, Derwent World Patents Index, vol. 1985, no. 45, Database accession no. 1985-279140, XP002717408 & JPS60188057 A 19850925 (KIKKOMAN CORP) [X] 1-5,8,11-13 * abstract *
 [XPY]  - DATABASE WPI, 0, Derwent World Patents Index, vol. 2009, no. 43, Database accession no. 2009-K99582, XP002717409 & JP2009095279 A 20090507 (DOKURITSU GYOSEI HOJIN NIPPON GAKUJUTSU) [XP] 1-5,8,11-13 * abstract * * paragraph [0035]; claim - * [Y] 1-13
 [Y]  - R. KOSZUL, "Stability of Large Segmental Duplications in the Yeast Genome", GENETICS, (20051215), vol. 172, no. 4, doi:10.1534/genetics.105.048058, ISSN 0016-6731, pages 2211 - 2222, XP055091584 [Y] 1-13 * the whole document *

DOI:   http://dx.doi.org/10.1534/genetics.105.048058
 [A]  - MATS E PETTERSSON ET AL, "Evolution of new gene functions: simulation and analysis of the amplification model", GENETICA ; AN INTERNATIONAL JOURNAL OF GENETICS AND EVOLUTION, KLUWER ACADEMIC PUBLISHERS, DO, (20080622), vol. 135, no. 3, ISSN 1573-6857, pages 309 - 324, XP019675960 [A] 1-13 * page 316, column 2 *
 [Y]  - PARAG Y ET AL, "GENETIC CONTROL OF CHROMOSOME INSTABILITY IN ASPERGILLUS-NIDULANS AS A MEAN FOR GENE AMPLIFICATION IN EUKARYOTIC MICROORGANISMS", MOLECULAR AND GENERAL GENETICS, (1975), vol. 140, no. 4, ISSN 0026-8925, pages 275 - 287, XP009174841 [Y] 1-13 * the whole document *
 [YP]  - DATABASE WPI, 0, Derwent World Patents Index, vol. 2010, no. 27, Database accession no. 2010-D91158, XP002717410 & JP2010075131 A 20100408 (YAMASA SHOYU KK) [YP] 1-13 * abstract *
International search[X]JPS57174087  (HIGETA SHOYU KK);
 [X]  - SMITH C. A. ET AL., "Silencing of the aflatoxin gene cluster in a diploid strain of Aspergillus flavus is suppressed by ectopic aflR expression.", GENETICS, (2007), vol. 176, no. 4, pages 2077 - 2086, XP008144795

DOI:   http://dx.doi.org/10.1534/genetics.107.073460
 [Y]  - KALAYANAMITR A. ET AL., "Occurrence of toxicity among protease, amylase, and color mutants of a nontoxic soy sauce koji mold.", APPL. ENVIRON. MICROBIOL., (1987), vol. 53, no. 8, pages 1980 - 1982, XP008144787
Examination   - KALAYANAMITR A ET AL, "Occurrence of toxicity among protease, amylase, and color mutants of a nontoxic soy sauce koji mold", APPLIED AND ENVIRONMENTAL MICROBIOLOGY, AMERICAN SOCIETY FOR MICROBIOLOGY, US, (19870801), vol. 53, no. 8, ISSN 0099-2240, pages 1980 - 1982, XP008144787
    - CARRIE A SMITH ET AL, "Silencing of the Aflatoxin Gene Cluster in a Diploid Strain of Aspergillus flavus Is Suppressed by Ectopic aflR Expression", GENE, GENETICS SOCIETY OF AMERICA, AUSTIN, TX, US, vol. 176, no. 4, doi:10.1534/GENETICS.107.073460, ISSN 0016-6731, (20070801), pages 2077 - 2086, (20070611), XP008144795

DOI:   http://dx.doi.org/10.1534/genetics.107.073460
by applicantJPH06153896
 JPH07274944
 JPH10210967
 JP2008054580
 JPH07222584
 JPH0970287
 JP2007228849
 JP2008306991
 JP2009100645
    - NATURE, (2005), vol. 438, pages 1157 - 61
    - FOOD SCI. TECHNOL. RES., (1999), vol. 5, no. 2, pages 153 - 155
    - DNA RES., (200808), vol. 15, no. 4, pages 172 - 83
    - Shoyu Shiken-Ho, JAPAN SOY SAUCE LABORATORY, (1985), page 287
    - Shoyu Shiken-Ho, JAPAN SOY SAUCE LABORATORY, (1985), page 104
The EPO accepts no responsibility for the accuracy of data originating from other authorities; in particular, it does not guarantee that it is complete, up to date or fit for specific purposes.