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Extract from the Register of European Patents

EP About this file: EP3045534

EP3045534 - FLUORESCENT PROTEIN [Right-click to bookmark this link]
StatusNo opposition filed within time limit
Status updated on  19.02.2021
Database last updated on 27.07.2024
FormerThe patent has been granted
Status updated on  13.03.2020
FormerGrant of patent is intended
Status updated on  29.11.2019
FormerExamination is in progress
Status updated on  29.10.2019
FormerGrant of patent is intended
Status updated on  22.08.2019
FormerExamination is in progress
Status updated on  08.12.2017
Most recent event   Tooltip08.07.2022Lapse of the patent in a contracting state
New state(s): MK		published on 10.08.2022  [2022/32]
Applicant(s)For all designated states
Osaka University
1-1 Yamadaoka
Suita-shi, Osaka 565-0871 / JP
[2016/29]
Inventor(s)01 / NAGAI, Takeharu
c/o Osaka University
1-1 Yamadaoka
Suita-shi Osaka 565-0871 / JP
02 / TIWARI, Dhermendra Kumar
c/o Osaka University
1-1 Yamadaoka
Suita-shi Osaka 565-0871 / JP
03 / ARAI, Yoshiyuki
c/o Osaka University
1-1 Yamadaoka
Suita-shi Osaka 565-0871 / JP
 [2016/29]
Representative(s)Dehns
St. Bride's House
10 Salisbury Square
London EC4Y 8JD / GB
[2020/16]
Former [2016/29]Wilkins, Christopher George
Dehns
St Bride's House
10 Salisbury Square
London EC4Y 8JD / GB
Application number, filing date14843850.011.09.2014
[2016/29]
WO2014JP74121
Priority number, dateJP2013019105813.09.2013         Original published format: JP 2013191058
[2016/29]
Filing languageJA
Procedural languageEN
PublicationType: A1 Application with search report
No.:WO2015037674
Date:19.03.2015
Language:JA
[2015/11]
Type: A1 Application with search report 
No.:EP3045534
Date:20.07.2016
Language:EN
[2016/29]
Type: B1 Patent specification 
No.:EP3045534
Date:15.04.2020
Language:EN
[2020/16]
Search report(s)International search report - published on:JP19.03.2015
(Supplementary) European search report - dispatched on:EP23.01.2017
ClassificationIPC:C12N15/09, C07K14/435, C07K19/00, C12N1/15, C12N1/19, C12N1/21, C12N5/10, G01N33/50, G01N21/64, C12Q1/02, C07K14/00, C07K14/47
[2017/08]
CPC:
C07K14/435 (EP,US); G01N21/6486 (US); C07K14/00 (US);
C07K14/47 (US); C07K14/4716 (US); C12Q1/02 (US);
G01N33/5005 (US); C07K2319/00 (EP,US); C07K2319/21 (US) (-)
Former IPC [2016/29]C12N15/09, C07K14/435, C07K19/00, C12N1/15, C12N1/19, C12N1/21, C12N5/10
Designated contracting statesAL,   AT,   BE,   BG,   CH,   CY,   CZ,   DE,   DK,   EE,   ES,   FI,   FR,   GB,   GR,   HR,   HU,   IE,   IS,   IT,   LI,   LT,   LU,   LV,   MC,   MK,   MT,   NL,   NO,   PL,   PT,   RO,   RS,   SE,   SI,   SK,   SM,   TR [2016/29]
Extension statesBANot yet paid
MENot yet paid
TitleGerman:FLUORESZENZPROTEIN[2016/29]
English:FLUORESCENT PROTEIN[2016/29]
French:PROTÉINE FLUORESCENTE[2016/29]
Entry into regional phase31.03.2016Translation filed 
31.03.2016National basic fee paid 
31.03.2016Search fee paid 
31.03.2016Designation fee(s) paid 
31.03.2016Examination fee paid 
Examination procedure31.03.2016Examination requested  [2016/29]
10.08.2017Amendment by applicant (claims and/or description)
12.12.2017Despatch of a communication from the examining division (Time limit: M04)
11.04.2018Reply to a communication from the examining division
04.06.2018Despatch of a communication from the examining division (Time limit: M04)
12.10.2018Reply to a communication from the examining division
14.11.2018Despatch of a communication from the examining division (Time limit: M04)
14.02.2019Reply to a communication from the examining division
23.08.2019Communication of intention to grant the patent
25.10.2019Disapproval of the communication of intention to grant the patent by the applicant or resumption of examination proceedings by the EPO
29.11.2019Communication of intention to grant the patent
03.03.2020Fee for grant paid
03.03.2020Fee for publishing/printing paid
03.03.2020Receipt of the translation of the claim(s)
Opposition(s)18.01.2021No opposition filed within time limit [2021/12]
Fees paidRenewal fee
19.09.2016Renewal fee patent year 03
19.09.2017Renewal fee patent year 04
18.09.2018Renewal fee patent year 05
18.09.2019Renewal fee patent year 06
Opt-out from the exclusive  Tooltip
competence of the Unified
Patent Court		 See the Register of the Unified Patent Court for opt-out data
Responsibility for the accuracy, completeness or quality of the data displayed under the link provided lies entirely with the Unified Patent Court.
Lapses during opposition  TooltipHU11.09.2014
AL15.04.2020
AT15.04.2020
CY15.04.2020
CZ15.04.2020
DK15.04.2020
EE15.04.2020
ES15.04.2020
FI15.04.2020
HR15.04.2020
IT15.04.2020
LT15.04.2020
LV15.04.2020
MC15.04.2020
MK15.04.2020
MT15.04.2020
NL15.04.2020
PL15.04.2020
RO15.04.2020
RS15.04.2020
SE15.04.2020
SI15.04.2020
SK15.04.2020
SM15.04.2020
TR15.04.2020
BG15.07.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
GB11.09.2020
IE11.09.2020
LU11.09.2020
BE30.09.2020
CH30.09.2020
FR30.09.2020
LI30.09.2020
[2022/32]
Former [2022/27]HU11.09.2014
AL15.04.2020
AT15.04.2020
CY15.04.2020
CZ15.04.2020
DK15.04.2020
EE15.04.2020
ES15.04.2020
FI15.04.2020
HR15.04.2020
IT15.04.2020
LT15.04.2020
LV15.04.2020
MC15.04.2020
MT15.04.2020
NL15.04.2020
PL15.04.2020
RO15.04.2020
RS15.04.2020
SE15.04.2020
SI15.04.2020
SK15.04.2020
SM15.04.2020
TR15.04.2020
BG15.07.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
GB11.09.2020
IE11.09.2020
LU11.09.2020
BE30.09.2020
CH30.09.2020
FR30.09.2020
LI30.09.2020
Former [2021/38]AL15.04.2020
AT15.04.2020
CZ15.04.2020
DK15.04.2020
EE15.04.2020
ES15.04.2020
FI15.04.2020
HR15.04.2020
IT15.04.2020
LT15.04.2020
LV15.04.2020
MC15.04.2020
NL15.04.2020
PL15.04.2020
RO15.04.2020
RS15.04.2020
SE15.04.2020
SI15.04.2020
SK15.04.2020
SM15.04.2020
BG15.07.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
GB11.09.2020
IE11.09.2020
LU11.09.2020
BE30.09.2020
CH30.09.2020
FR30.09.2020
LI30.09.2020
Former [2021/37]AL15.04.2020
AT15.04.2020
CZ15.04.2020
DK15.04.2020
EE15.04.2020
ES15.04.2020
FI15.04.2020
HR15.04.2020
IT15.04.2020
LT15.04.2020
LV15.04.2020
MC15.04.2020
NL15.04.2020
PL15.04.2020
RO15.04.2020
RS15.04.2020
SE15.04.2020
SI15.04.2020
SK15.04.2020
SM15.04.2020
BG15.07.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
IE11.09.2020
LU11.09.2020
BE30.09.2020
CH30.09.2020
FR30.09.2020
LI30.09.2020
Former [2021/36]AL15.04.2020
AT15.04.2020
CZ15.04.2020
DK15.04.2020
EE15.04.2020
ES15.04.2020
FI15.04.2020
HR15.04.2020
IT15.04.2020
LT15.04.2020
LV15.04.2020
MC15.04.2020
NL15.04.2020
PL15.04.2020
RO15.04.2020
RS15.04.2020
SE15.04.2020
SI15.04.2020
SK15.04.2020
SM15.04.2020
BG15.07.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
LU11.09.2020
BE30.09.2020
FR30.09.2020
Former [2021/35]AL15.04.2020
AT15.04.2020
CZ15.04.2020
DK15.04.2020
EE15.04.2020
ES15.04.2020
FI15.04.2020
HR15.04.2020
IT15.04.2020
LT15.04.2020
LV15.04.2020
MC15.04.2020
NL15.04.2020
PL15.04.2020
RO15.04.2020
RS15.04.2020
SE15.04.2020
SI15.04.2020
SK15.04.2020
SM15.04.2020
BG15.07.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
LU11.09.2020
FR30.09.2020
Former [2021/27]AL15.04.2020
AT15.04.2020
CZ15.04.2020
DK15.04.2020
EE15.04.2020
ES15.04.2020
FI15.04.2020
HR15.04.2020
IT15.04.2020
LT15.04.2020
LV15.04.2020
MC15.04.2020
NL15.04.2020
PL15.04.2020
RO15.04.2020
RS15.04.2020
SE15.04.2020
SI15.04.2020
SK15.04.2020
SM15.04.2020
BG15.07.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
LU11.09.2020
Former [2021/23]AL15.04.2020
AT15.04.2020
CZ15.04.2020
DK15.04.2020
EE15.04.2020
ES15.04.2020
FI15.04.2020
HR15.04.2020
IT15.04.2020
LT15.04.2020
LV15.04.2020
MC15.04.2020
NL15.04.2020
PL15.04.2020
RO15.04.2020
RS15.04.2020
SE15.04.2020
SI15.04.2020
SK15.04.2020
SM15.04.2020
BG15.07.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
Former [2021/10]AL15.04.2020
AT15.04.2020
CZ15.04.2020
DK15.04.2020
EE15.04.2020
ES15.04.2020
FI15.04.2020
HR15.04.2020
IT15.04.2020
LT15.04.2020
LV15.04.2020
NL15.04.2020
PL15.04.2020
RO15.04.2020
RS15.04.2020
SE15.04.2020
SK15.04.2020
SM15.04.2020
BG15.07.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
Former [2021/09]AL15.04.2020
AT15.04.2020
CZ15.04.2020
DK15.04.2020
EE15.04.2020
ES15.04.2020
FI15.04.2020
HR15.04.2020
IT15.04.2020
LT15.04.2020
LV15.04.2020
NL15.04.2020
RO15.04.2020
RS15.04.2020
SE15.04.2020
SM15.04.2020
BG15.07.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
Former [2021/08]AL15.04.2020
DK15.04.2020
ES15.04.2020
FI15.04.2020
HR15.04.2020
LT15.04.2020
LV15.04.2020
NL15.04.2020
RO15.04.2020
RS15.04.2020
SE15.04.2020
SM15.04.2020
BG15.07.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
Former [2021/04]AL15.04.2020
FI15.04.2020
HR15.04.2020
LT15.04.2020
LV15.04.2020
NL15.04.2020
RS15.04.2020
SE15.04.2020
BG15.07.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
Former [2021/01]FI15.04.2020
HR15.04.2020
LT15.04.2020
LV15.04.2020
NL15.04.2020
RS15.04.2020
SE15.04.2020
BG15.07.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
Former [2020/51]FI15.04.2020
HR15.04.2020
LT15.04.2020
LV15.04.2020
NL15.04.2020
RS15.04.2020
SE15.04.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
Former [2020/50]FI15.04.2020
HR15.04.2020
LT15.04.2020
LV15.04.2020
NL15.04.2020
SE15.04.2020
NO15.07.2020
GR16.07.2020
IS15.08.2020
PT17.08.2020
Former [2020/48]FI15.04.2020
LT15.04.2020
NL15.04.2020
SE15.04.2020
NO15.07.2020
IS15.08.2020
PT17.08.2020
Former [2020/47]LT15.04.2020
NO15.07.2020
IS15.08.2020
Former [2020/46]IS15.08.2020
Documents cited:Search[A]  - SUSAN GAYDA ET AL, "Mechanistic Insights into Reversible Photoactivation in Proteins of the GFP Family", BIOPHYSICAL JOURNAL, AMSTERDAM, NL, (20121201), vol. 103, no. 12, doi:10.1016/j.bpj.2012.11.011, ISSN 0006-3495, pages 2521 - 2531, XP055334155 [A] 1-10 * the whole document *

DOI:   http://dx.doi.org/10.1016/j.bpj.2012.11.011
International search[Y]WO2005113772  (RIKEN [JP], et al);
 [X]  - ANDRESEN, M. ET AL., "Photoswitchable fluorescent proteins enable monochromatic multilabel imaging and dual color fluorescence nanoscopy", NAT. BIOTECHNOL., (2008), vol. 26, no. 9, pages 1035 - 1040, XP055015979

DOI:   http://dx.doi.org/10.1038/nbt.1493
 [X]  - BRAKEMANN, T. ET AL., "Molecular Basis of the Light-driven Switching of the Photochromic Fluorescent Protein Padron", J. BIOL. CHEM., (2010), vol. 285, no. 19, pages 14603 - 14609, XP055327180

DOI:   http://dx.doi.org/10.1074/jbc.M109.086314
by applicantWO2005113772
    - TANJA BRAKEMANN ET AL., J. BIOL. CHEM., (2010), vol. 285, pages 14603 - 14609
    - GAYDA, S.; NIENHAUS, K.; NIENHAUS, G.U., BIOPHYSICAL JOURNAL, (2012), vol. 103, pages 2521 - 31
The EPO accepts no responsibility for the accuracy of data originating from other authorities; in particular, it does not guarantee that it is complete, up to date or fit for specific purposes.