blank Quick help
blank Maintenance news

Scheduled maintenance

Regular maintenance outages:
between 05.00 and 05.15 hrs CET (Monday to Sunday).

Other outages
Availability
Register Forum

2022.02.11

More...
blank News flashes

News Flashes

New version of the European Patent Register – SPC proceedings information in the Unitary Patent Register.

2024-07-24

More...
blank Related links

Extract from the Register of European Patents

EP About this file: EP3976633

EP3976633 - GENE FOR PARTHENOGENESIS [Right-click to bookmark this link]
StatusRequest for examination was made
Status updated on  04.03.2022
Database last updated on 31.08.2024
FormerThe international publication has been made
Status updated on  05.12.2020
Formerunknown
Status updated on  08.06.2020
Most recent event   Tooltip22.05.2024New entry: Renewal fee paid 
Applicant(s)For all designated states
Keygene N.V.
P.O. Box 216
6700 AE Wageningen / NL
[2022/14]
Inventor(s)01 / UNDERWOOD, Charles Joseph
c/o P.O. Box 216
6700 AE Wageningen / NL
02 / RIGOLA, Diana
c/o P.O. Box 216
6700 AE Wageningen / NL
03 / VAN DIJK, Peter Johannes
c/o P.O. Box 216
6700 AE Wageningen / NL
04 / OP DEN CAMP, Rik Hubertus Martinus
c/o P.O. Box 216
6700 AE Wageningen / NL
05 / SCHRANZ, Michael, Eric
c/o Droevendaalsesteeg 4
6708 PB Wageningen / NL
06 / VIJVERBERG, Catharina, Adriana
c/o Droevendaalsesteeg 4
6708 PB Wageningen / NL
 [2022/14]
Representative(s)Nederlandsch Octrooibureau
P.O. Box 29720
2502 LS The Hague / NL
[2022/14]
Application number, filing date20729068.529.05.2020
[2022/14]
WO2020EP64991
Priority number, dateEP2019017725229.05.2019         Original published format: EP 19177252
EP2019020087201.10.2019         Original published format: EP 19200872
EP2020017024317.04.2020         Original published format: EP 20170243
[2022/14]
Filing languageEN
Procedural languageEN
PublicationType: A1 Application with search report
No.:WO2020239984
Date:03.12.2020
Language:EN
[2020/49]
Type: A1 Application with search report 
No.:EP3976633
Date:06.04.2022
Language:EN
The application published by WIPO in one of the EPO official languages on 03.12.2020 takes the place of the publication of the European patent application.
[2022/14]
Search report(s)International search report - published on:EP03.12.2020
ClassificationIPC:C07K14/415, C12N15/82
[2022/14]
CPC:
C07K14/415 (EP,IL,US); C12N15/8287 (EP,IL,US)
Designated contracting statesAL,   AT,   BE,   BG,   CH,   CY,   CZ,   DE,   DK,   EE,   ES,   FI,   FR,   GB,   GR,   HR,   HU,   IE,   IS,   IT,   LI,   LT,   LU,   LV,   MC,   MK,   MT,   NL,   NO,   PL,   PT,   RO,   RS,   SE,   SI,   SK,   SM,   TR [2022/14]
TitleGerman:GEN FÜR PARTHOGENESE[2022/14]
English:GENE FOR PARTHENOGENESIS[2022/14]
French:GÈNE POUR PARTHÉNOGENÈSE[2022/14]
Entry into regional phase14.12.2021National basic fee paid 
14.12.2021Designation fee(s) paid 
14.12.2021Examination fee paid 
Examination procedure14.12.2021Examination requested  [2022/14]
14.12.2021Date on which the examining division has become responsible
28.07.2022Amendment by applicant (claims and/or description)
Fees paidRenewal fee
30.05.2022Renewal fee patent year 03
30.05.2023Renewal fee patent year 04
22.05.2024Renewal fee patent year 05
Opt-out from the exclusive  Tooltip
competence of the Unified
Patent Court
See the Register of the Unified Patent Court for opt-out data
Responsibility for the accuracy, completeness or quality of the data displayed under the link provided lies entirely with the Unified Patent Court.
Cited inInternational search[Y]EP2530160  (LEIBNIZ INST FUER PFLANZENGENETIK UND KULTURPFLANZENFORSCHUNG GATERSLEBEN IPK [DE]) [Y] 1-15 * claims 1-19 *;
 [Y]WO2015061355  (UNIV GEORGIA [US], et al) [Y] 1-15 * claims 1-28 *;
 [YD]WO2017039452  (KEYGENE NV [NL]) [YD] 1-15 * claims 1-48 *;
 [A]  - KITTY VIJVERBERG ET AL, "Identifying and Engineering Genes for Parthenogenesis in Plants", FRONTIERS IN PLANT SCIENCE, CH, (20190219), vol. 10, doi:10.3389/fpls.2019.00128, ISSN 1664-462X, XP055633366 [A] 1-15 * the whole document *

DOI:   http://dx.doi.org/10.3389/fpls.2019.00128
by applicantEP0067553
 US4407956
 WO8402913
 EP0116718
 EP0120561
 EP0120515
 WO8501856
 US4536475
 EP0223247
 US4684611
 EP0242246
 EP0242236
 EP0270356
 EP0270822
 EP0342926
 WO9006999
 EP0426641
 EP0507698
 EP0508909
 US5164316
 EP0534858
 US5254799
 WO9500555
 WO9506722
 EP0686191
 WO9606932
 US5510471
 US5527695
 US5591616
 WO9710704
 US5635618
 US5641876
 WO9737012
 US5693507
 WO9748819
 EP0506763
 US6031151
 US6051753
 WO0026371
 US6063985
 WO0056897
 WO0141558
 US2002138879
 US6563026
 US2004148667
 US2004168216
 US2005155111
 US2006179498
 WO2007000067
 WO2007066214
 WO2017039452
    - HENIKOFFHENIKOFF, PNAS, (19920000), vol. 89, pages 915 - 919
    - ALTSCHUL et al., J. Mol. Biol., (19900000), vol. 215, pages 403 - 10
    - ALTSCHUL et al., Nucleic Acids Res., (19970000), vol. 25, no. 17, pages 3389 - 3402
    - GUO et al., Scientific Reports, (20170601), vol. 7, no. 1, page 2634
    - BENNETZEN J.L.HALL B.D, J. Biol. Chem., (19820000), vol. 257, pages 3026 - 3031
    - ITAKURA et al., Science, (19770000), vol. 198, pages 1056 - 1063
    - CORNEJO et al., Plant Mol.Biol., (19930000), vol. 23, pages 567 - 581
    - NAKAMURA et al., Nucl. Acids Res., (20000000), vol. 28, page 292
    - SPRUNCK et al., Science, (20120000), vol. 338.6110, pages 1093 - 1097
    - STEFFEN et al., Plant Journal, (20070000), vol. 51, pages 281 - 292
    - OHNISHI et al., PlantPhysiology, (20140000), vol. 165, pages 1533 - 1543
    - MCCALLUM et al., Nat Biotech, (20000000), vol. 18, page 455
    - MCCALLUM et al., Plant Physiol., (20000000), vol. 123, pages 439 - 442
    - AN et al., Plant J., (19960000), vol. 10, page 107
    - AOYAMACHUA, Plant Journal, (19970000), vol. 11, pages 605 - 612
    - ASKER, S., "Progress in apomixis research", Hereditas, (19790000), vol. 91, no. 2, pages 231 - 240
    - LAST et al., Theor. Appl. Genet., (19900000), vol. 81, pages 581 - 588
    - AUSUBEL et al., "Current Protocols in Molecular Biology", Current Protocols, (19940000), vol. 1 and 2
    - BAE T.W.PARK R.H.KWAK Y.S.LEE H.Y.RYU S.B., "Agrobacterium tumefaciens-mediated transformation of a medicinal plant Taraxacum platycarpum", Plant Cell, Tissue and Organ Culture, (20050000), vol. 80, pages 50 - 57
    - BAULCOMBE D.C., Plant Mol Biol., (19961000), vol. 32, no. 1-2, pages 79 - 88
    - BARRELLGROSSNIKLAUS, "Confocal microscopy of whole ovules for analysis of reproductive development: the elongate1 mutant affects meiosis II", Plant Journal, (20050000), vol. 34, doi:10.1111/j.1365-313X.2005.02456.x, pages 309 - 320, XP055004256

DOI:   http://dx.doi.org/10.1111/j.1365-313X.2005.02456.x
    - BICKNELLKOLTUNOW, "Understanding apomixis: recent advances and remaining conundrums", The Plant Cell, (20040000), vol. 16, doi:10.1105/TPC.017921, pages S228 - S245, XP002585046

DOI:   http://dx.doi.org/10.1105/TPC.017921
    - BIH et al., J. Biol. Chem., (19990000), vol. 274, pages 22884 - 22894
    - BOREVITZ, J.O.LIANG, D.PLOUFFE, D.CHANG, H.-S.ZHU, T.WEIGEL, D.BERRY, C.C.WINZELER, E.CHORY, J., "Large-scale identification of single-feature polymorphisms in Arabidopsis", Genome Res., (20030000), vol. 13, pages 513 - 523
    - BORTESI, L.FISCHER, R., "The CRISPR/Cas9 system for plant genome editing and beyond", Biotechnology Advanced, (20150000), vol. 33, no. 1, doi:10.1016/j.biotechadv.2014.12.006, pages 41 - 52, XP055566672

DOI:   http://dx.doi.org/10.1016/j.biotechadv.2014.12.006
    - BRUCE MHESS ABAI JMAULEON RDIAZ M GSUGIYAMA NBORDEOS AWANG GLEUNG HLEACH, J., "Detection of genomic deletions in rice using oligonucleotide microarrays", BMC Genomics, (20090000), vol. 10, pages 129 - 140
    - CATANACH ASERASMUSON SKPODIVINSKY EJORDAN BRBICKNELL R., "Deletion mapping of genetic regions associated with apomixis in Hieracium", Proc. Nat. Acad. Sci., (20060000), vol. 103, doi:10.1073/PNAS.0605588103, pages 18650 - 5, XP002583195

DOI:   http://dx.doi.org/10.1073/PNAS.0605588103
    - CHRISTENSEN et al., Plant Mol. Biol., (19920000), vol. 18, pages 675 - 689
    - CHUPEAU et al., "Transgenic plants of lettuce (Lactuca sativa) obtained through electroporation of protoplasts", Bio/Technology, (19890000), vol. 7, pages 503 - 508
    - CORDERA et al., The Plant Journal, (19940000), vol. 6, page 141
    - CORNELISSEN et al., EMBO J., (19860000), vol. 5, pages 37 - 40
    - CRISMANI W. et al., J. Exp. Bot., (20130000), vol. 64, pages 55 - 65
    - CURTIS IS et al., J. Exp. Bot., (19940000), vol. 45.10, pages 1441 - 1449
    - DANIELL, H., "Molecular strategies for gene containment in transgenic crops", Nature biotechnology, (20020000), vol. 20, doi:10.1038/nbt0602-581, pages 581 - 586, XP037103842

DOI:   http://dx.doi.org/10.1038/nbt0602-581
    - DE PATER et al., Plant J., (19920000), vol. 2, pages 834 - 844
    - DEPICKER A.VAN MONTAGU M., "Post-transcriptional gene silencing in plants", Current Opinion in Cell Biology, (19970000), vol. 9, pages 373 - 382
    - DEPICKER et al., J. Mol. Appl. Genetics, (19820000), vol. 1, pages 561 - 573
    - ENGLBRECHT et al., BMC Genomics, (20040000), vol. 5, no. 1, page 39
    - VIELLE-CALZADA, J-PH.B.L. BURSONE.C BASHAWM. A. HUSSEY, "Early fertilization events in the sexual an aposporous egg apparatus of Pennisetum ciliare (L.) Link", The Plant Journal, (19950000), vol. 8, no. 2, pages 309 - 316
    - LIU et al., Genomics, (19950000), vol. 25, no. 3, pages 674 - 81
    - FLOREZ-RUEDA et al., "Laser-Assisted Microdissection of Plant Embryos for Transcriptional Profiling", Methods Mol Biol, (20200000), vol. 2122, pages 127 - 139
    - FRANCK et al., Cell, (19800000), vol. 21, pages 285 - 294
    - GARDNER et al., Nucleic Acids Research, (19810000), vol. 9, pages 2871 - 2887
    - GATZ, Annu Rev Plant Physiol Plant Mol Biol., (19970000), vol. 48, pages 89 - 108
    - VELTEN et al., EMBO J, (19840000), vol. 3, pages 2723 - 2730
    - GUO et al., Scientific reports, (20170601), vol. 7, no. 1, page 2634
    - GOULD et al., Plant Physiol., (19910000), vol. 95, pages 426 - 434
    - GRIMANELLI D., Curr. Opin. Plant Biol., (20120000), vol. 15, pages 57 - 62
    - HASHIMSHONY, T.SENDEROVICH, N.AVITAL, G. et al., "CEL-Seq2: sensitive highly-multiplexed single-cell RNA-Seq", Genome Biol, (20160000), vol. 17, page 77
    - HASHIMSHONY TWAGNER FSHER NYANAI I., "CEL-Seq: single-cell RNA-Seq by multiplexed linear amplification", Cell Rep., (20120000), vol. 2, no. 3, doi:10.1016/j.celrep.2012.08.003, pages 666 - 673, XP055111758

DOI:   http://dx.doi.org/10.1016/j.celrep.2012.08.003
    - HELLIWELLWATERHOUSE, Methods, (20030000), vol. 30, no. 4, pages 289 - 95
    - HERMSEN, J. G. TH., "Breeding for apomixis in potato: Pursuing a utopian scheme", Euphytica, (19800000), vol. 29, pages 595 - 607
    - HESSE et al., EMBO J., (19890000), vol. 8, pages 2453 - 2461
    - HOLMES, M, Historical Studies in the Natural Sciences, (20180000), vol. 48, no. 1, ISSN 1939-1811, pages 1 - 23
    - HULLHOWELL, Virology, (19870000), vol. 86, pages 482 - 493
    - KAGALE et al., Plant Physiology, (20100000), vol. 152, pages 1009 - 1134
    - KEIL et al., Nucl. Acids Res., (19860000), vol. 14, pages 5641 - 5650
    - KLOSGENWEIL, Mol. Gen. Genet., (19910000), vol. 225, pages 297 - 304
    - KLOSGEN et al., Mol. Gen. Genet., (19890000), vol. 217, pages 155 - 161
    - LIU et al., Methods Mol. Biol., (20050000), vol. 286, pages 341 - 8
    - LOVE et al., Plant J., (20000000), vol. 21, pages 579 - 88
    - LUTZ KA et al., Plant J., (20040000), vol. 37, no. 6, pages 906 - 13
    - MAILLON et al., FEMS Microbiol. Letters, (19890000), vol. 60, pages 205 - 210
    - MAXINGLIANG et al., "A robust CRISPR/Cas9 system for convenient, high-efficiency multiplex genome editing in monocot and dicot plants", Molecular plant, (20150000), vol. 8.8, pages 1274 - 1284
    - MC BRIDE et al., Bio/Technology, (19950000), vol. 13, page 362
    - MICHELMORE, R.W.MARSH, E.SEELY, S.LANDRY, B., "Transformation of lettuce (Lactuca sativa) mediated by Agrobacterium tumefaciens", Plant Cell Rep., (19870000), vol. 6, pages 439 - 442
    - MICHELMORE, R.W.PARAN, I.KESSELI, R.V., "Identification of markers linked to disease resistance genes by bulked segregant analysis: a rapid method to detect markers in specific genomic regions using segregating populations", Proc. Natl. Acad. Sci., (19910000), vol. 88, doi:10.1073/pnas.88.21.9828, pages 9828 - 9832, XP002184162

DOI:   http://dx.doi.org/10.1073/pnas.88.21.9828
    - MORGAN, R.OZIAS-AKINS, P.HANNA, W.W., "Seed set in an apomictic BC3 pearl millet", Int. J. Plant Sci., (19980000), vol. 159, pages 89 - 97
    - MORRIS et al., Biochem. Biophys. Res. Commun., (19990000), vol. 255, pages 328 - 333
    - MULLER, K.J.HE, X.FISCHER, R.PRUFER, D., "Constitutive knox1 gene expression in dandelion (Taraxacum officinale, Web.) changes leaf morphology from simple to compound", Planta, (20060000), vol. 224, doi:10.1007/s00425-006-0288-y, pages 1023 - 1027, XP019443306

DOI:   http://dx.doi.org/10.1007/s00425-006-0288-y
    - NEKRASOVVLADIMIR et al., "Targeted mutagenesis in the model plant Nicotiana benthamiana using Cas9 RNA-guided endonuclease", Nature biotechnology, (20130000), vol. 31.8, page 691
    - VERDAGUER et al., Plant Mol. Biol., (19980000), vol. 37, pages 1055 - 1067
    - VAN DEN BROECK et al., Nature, (19850000), vol. 313, pages 358 - 812
    - OELMULLER et al., Mol. Gen. Genet., (19930000), vol. 237, pages 261 - 272
    - OZIAS-AKINS, P.P.J. VAN DIJK., "Mendelian genetics of apomixis in plants", Annu. Rev. Genet., (20070000), vol. 41, doi:10.1146/ANNUREV.GENET.40.110405.090511, pages 509 - 537, XP002583194

DOI:   http://dx.doi.org/10.1146/ANNUREV.GENET.40.110405.090511
    - PARK et al., J.Biol. Chem., (19970000), vol. 272, pages 6876 - 6881
    - RIOS GNARANJO M AIGLESIAS D JRUIZ-RIVERO OGERAUD, MUSACH, ATALON M., "Characterization of hemizygous deletions in Citrus using array-Comparative Genomic Hybridization and microsynteny comparisons with the poplar genome", BMC Genomics, (20080000), vol. 9, pages 381 - 395
    - "Transfer of apomixis through wide crosses", SAVIDAN Y., The flowering of apomixis: From mechanisms to genetic engineering, CIMMYT, IRD, (20010000), pages 153 - 167
    - SHCHERBAN et al., Proc. Natl. Acad. Sci USA, (19950000), vol. 92, pages 9245 - 9249
    - SIDOROV VA et al., Plant J., (19990000), vol. 19, pages 209 - 216
    - SMITH TFWATERMAN MS, J. Mol. Biol, (19810000), vol. 147, no. 1, pages 195 - 7
    - STAM, M.MOL, J.N.KOOTER, J.M., "The silencing of genes in transgenic plants", Annals of Botany, (19970000), vol. 79, pages 3 - 12
    - SUTLIFF et al., Plant Molec. Biol., (19910000), vol. 16, pages 579 - 591
    - TAS, I.C.Q.VAN DIJK, P.J., "Crosses between sexual and apomictic dandelions (Taraxacum) I. The inheritance of apomixis", Heredity, (19990000), vol. 83, doi:10.1046/j.1365-2540.1999.00619.x, pages 707 - 714, XP002739182

DOI:   http://dx.doi.org/10.1046/j.1365-2540.1999.00619.x
    - TAVLADORAKI et al., FEBS Lett., (19980000), vol. 426, pages 62 - 66
    - TERASHIMA et al., Appl. Microbiol. Biotechnol., (19990000), vol. 52, pages 516 - 523
    - VAECK et al., Nature, (19870000), vol. 328, pages 33 - 37
    - VAN BAARLENDE JONG, J.H.VAN DIJK, P.J., "Comparative cyto-embryological investigations of sexual and apomictic dandelions (Taraxacum) and their apomictic hybrids", Sex Plant Reprod, (20020000), vol. 15, pages 31 - 38
    - VAN DIJK, P.J.BAKX-SCHOTMAN, J.M.T., "Formation of unreduced megaspores (diplospory) in apomictic dandelions (Taraxacum) is controlled by a sex-specific dominant gene", Genetics, (20040000), vol. 166, doi:10.1534/genetics.166.1.483, pages 483 - 492, XP009073544

DOI:   http://dx.doi.org/10.1534/genetics.166.1.483
    - VAN DIJK, P.J.SCHAUER, S.E.VELTENSCHELL, Nucleic Acids Research, vol. 13, (19850000), pages 6981 - 6998, URL: https://www.keygene.com/wp-content/uploads/2018/07/apomixis-game-changer-in-breeding.pdf
    - VAN DIJK, P.J.RIGOLA, D.SCHAUER, S.E., "Plant breeding: surprisingly, less sex is better", Current Biology, (20160000), vol. 26.3, pages R122 - R124
    - VAN DIJK, P.J.TAS, I.C.Q.FALQUE, MBAKX-SCHOTMAN J.M.T., "Crosses between sexual and apomictic dandelions (Taraxacum). II. The breakdown of apomixis", Heredity, (19990000), vol. 83, pages 715 - 721
    - VAN DIJK, P.J.VAN BAARLEN, P.DE JONG, J.H., "The occurrence of phenotypically complementary apomixis-recombinants in crosses between sexual and apomictic dandelions (Taraxacum officinale", Sex. Plant Repr., (20030000), vol. 16, pages 71 - 76
    - VIELLE-CALZADA, J.P.CRANE, C.F.STELLY, D.M., "Apomixis: The asexual revolution", Science, (19960000), vol. 274, pages 1322 - 1323
    - VIJVERBERG, K.VAN DER HULST, R.LINDHOUT, P.VAN DIJK P.J., "A genetic linkage map of the diplosporous chromosomal region in Taraxacum (common dandelion; Asteraceae", Theor. Appl. Genet., (20040000), vol. 108, doi:10.1007/s00122-003-1474-y, pages 725 - 732, XP055320376

DOI:   http://dx.doi.org/10.1007/s00122-003-1474-y
    - VOS, P.HOGERS, R.BLEEKER, M.REIJANS, M.LEE, TH.VAN DER, HOMES, M.FRIJTERS, A.POT, J.PELEMAN, J.KUIPER, M., "AFLP: a new technique for DNA fingerprinting", Nucl. Acids Res., (19950000), vol. 23, pages 4407 - 4414
    - WESLEY et al., Methods Mol Biol., (20030000), vol. 236, pages 273 - 86
    - WESLEY et al., Methods Mol Biol., (20040000), vol. 265, pages 117 - 30
    - WONG et al., Plant Molec. Biol., (19920000), vol. 20, pages 81 - 93
    - WU KK1BURNQUIST WSORRELLS METEW TLMOORE PHTANKSLEY SD, "The detection and estimation of linkage in polyploids using single-dose restriction fragments", Theor. Appl. Genet., (19920000), vol. 83, pages 294 - 300
    - WUEST SEVIJVERBERG KSCHMIDT A et al., "Arabidopsis female gametophyte gene expression map reveals similarities between plant and animal gametes", Curr Biol., (20100000), vol. 20, no. 6, pages 506 - 512, XP026977654
    - ZHANG et al., The Plant Cell, (19910000), vol. 3, pages 1155 - 1165
The EPO accepts no responsibility for the accuracy of data originating from other authorities; in particular, it does not guarantee that it is complete, up to date or fit for specific purposes.